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Species delimitation and taxonomic revision of Oxyopes (Araneae: Oxyopidae) of Taiwan, with description of two new species
- 1. Department of Life Science, National Taiwan Normal University, No. 88, Sec. 4, Tingzhou Rd., Taipei 11677, Taiwan & Zoology Division, Taiwan Endemic Species Research Institute, No. 1, Minsheng E. Rd., Jiji Township, Nantou 55244, Taiwan. 80443007s@gapps.ntnu.edu.tw; https://orcid.org/0000-0002-2862-5155
- 2. Department of Life Sciences, National Chung Hsing University, No. 145, Xingda Rd., South Dist., Taichung 40227, Taiwan. bolasargiope@email.nchu.edu.tw; https://orcid.org/0000-0001-9785-718X & Research Center for Global Change Biology, National Chung Hsing University, No. 145, Xingda Rd., South Dist., Taichung 40227, Taiwan
- 3. Department of Life Science, National Taiwan Normal University, No. 88, Sec. 4, Tingzhou Rd., Taipei 11677, Taiwan
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Lo, Ying-Yuan, Cheng, Ren-Chung, Lin, Chung-Ping (2021): Species delimitation and taxonomic revision of Oxyopes (Araneae: Oxyopidae) of Taiwan, with description of two new species. Zootaxa 4927 (1): 58-86, DOI: 10.11646/zootaxa.4927.1.4
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References
- Agnarsson, I., LeQuier, S.M., Kuntner, M., Cheng, R.-C., Coddington, J.A. & Binford, G. (2016) Phylogeography of a good Caribbean disperser: Argiope argentata (Araneae, Araneidae) and a new "cryptic" species from Cuba. ZooKeys, 625, 25-44. https://doi.org/10.3897/zookeys.625.8729
- Baehr, B.C., Harms, D., Duperre, N. & Raven, R. (2017) The Australian Lynx Spiders (Araneae, Oxyopidae, Oxyopes) of the Godeffroy Collection, including the description of a new species. Evolutionary Systematics, 1, 11-37. https://doi. org/10.3897/evolsyst.1.14652
- Barrett, R.D.H. & Hebert, P.D.N. (2005) Identifying spiders through DNA barcodes. Canadian Journal of Zoology, 83 (3), 481-491. https://doi.org/10.1139/z05-024
- Basnet, K. & Mukhopadhyay, A. (2014) Biocontrol potential of the lynx spider Oxyopes javanus (Araneae: Oxyopidae) against the tea mosquito bug, Helopeltis theivora (Heteroptera: Miridae). International Journal of Tropical Insect Science, 34, 232-238. https://doi.org/10.1017/S1742758414000538
- Bickford, D., Lohman, D.J., Sodhi, N.S., Ng, P.K.L., Meier, R., Winker, K., Ingram, K.K. & Das, I. (2007) Cryptic species as a window on diversity and conservation. Trends in Ecology & Evolution, 22, 148-155. https://doi.org/10.1016/j.tree.2006.11.004
- Blagoev, G.A., deWaard, J.R., Ratnasingham, S., deWaard, S.L., Lu, L., Robertson, J., Telfer A.C. & Hebert, P.D.N. (2016) Untangling taxonomy: a DNA barcode reference library for Canadian spiders. Molecular Ecology Resources, 16, 325-341. https://doi.org/10.1111/1755-0998.12444
- Bˆ senberg, W. & Strand, E. (1906) Japanische Spinnen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 30, 93-422.
- Bouckaert, R., Heled, J., Kuhnert, D., Vaughan, T., Wu, C.H., Xie, D., Suchard, M.A., Rambaut, A. & Drummond, A.J. (2014) BEAST 2: A Software Platform for Bayesian Evolutionary Analysis. PLoS Computational Biology, 10 (4), e1003537. https://doi.org/10.1371/journal.pcbi.1003537
- Brady, A.R. (1964) The lynx spiders of North America, north of Mexico (Araneae: Oxyopidae). Bulletin of the Museum of Comparative Zoology, 131, 429-518.
- Brady, A.R. & Santos, A.J. (2005) Oxyopidae. In: Ubick, D., Paquin, P., Cushing, P.E. & Roth, V. (Eds.), Spiders of North America: an identification manual. American Arachnological Society, Keene, New Hampshire, pp. 189-191.
- Brown, S.D.J., Collins, R.A., Boyer, S., Lefort, M.C., Malumbres-Olarte, J., Vink, C.J. & Cruickshank, R.H. (2012) SPIDER: an R package for the analysis of species identity and evolution, with particular reference to DNA barcoding. Molecular Ecology Resources, 12, 562-565. https://doi.org/10.1111/j.1755-0998.2011.03108.x
- Candek, K. & Kuntner, M. (2015) DNA barcoding gap: reliable species identification over morphological and geographical scales. Molecular Ecology Resources, 15, 268-277. https://doi.org/10.1111/1755-0998.12304
- Cao, X., Liu, J., Chen, J., Zheng, G., Kuntner, M. & Agnarsson, I. (2016) Rapid dissemination of taxonomic discoveries based on DNA barcoding and morphology. Scientific Reports, 6, 37066. https://doi.org/10.1038/srep37066
- Chamberlin, R.V. (1924) Descriptions of new American and Chinese spiders, with notes on other Chinese species. Proceedings of the United States National Museum, 63 (13), 1-38. https://doi.org/10.5479/si.00963801.63-2481.1
- Chen, Y.T., Tseng, H.Y., Jeng, M.L., Su, Y.C., Huang, W.S. & Lin, C.P. (2017) Integrated species delimitation and conservation implications of an endangered weevil Pachyrhynchus sonani (Coleoptera: Curculionidae) in Green and Orchid Islands of Taiwan. Systematic Entomology, 42, 796-813. https://doi.org/10.1111/syen.12242
- Chu, Y.I. & Okuma, C. (1970) Preliminary survey on the spider-fauna of the paddy fields in Taiwan. Mushi, 44, 65-88.
- Crews, S.C. & Harvey, M.S. (2011) The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. Zookeys, 99, 1-103. https://doi.org/10.3897/zookeys.99.723
- Darriba, D., Taboada, G.L., Doallo, R. & Posada, D. (2012) JModelTest 2: More Models, New Heuristics and Parallel Computing. Nature Methods, 9 (8), 772-772. https://doi.org/10.1038/nmeth.2109
- Deeleman-Reinhold, C.L. (2009) Description of the lynx spiders of a canopy fogging project in northern Borneo (Araneae: Oxyopidae), with description of a new genus and six new species of Hamataliwa. Zoologische Mededelingen, 83, 673-700.
- Drummond, A.J. & Rambaut, A. (2007) BEAST: Bayesian Evolutionary Analysis by Sampling Trees. BMC Evolutionary Biology, 7 (1), 214. https://doi.org/10.1186/1471-2148-7-214
- Duncan, R.P., Rynerson, M.R., Ribera, C. & Binford, G.J. (2010) Diversity of Loxosceles spiders in Northwestern Africa and molecular support for cryptic species in the Loxosceles rufescens lineage. Molecular Phylogenetics and Evolution, 55, 234-248. https://doi.org/10.1016/j.ympev.2009.11.026
- Esselstyn, J.A., Evans, B.J., Sedlock, J.L., Anwarali Khan, F.A. & Heaney, L.R. (2012) Single-locus species delimitation: a test of the mixed Yule-coalescent model, with an empirical application to Philippine round-leaf bats. Proceedings of the Royal Society B, 279, 3678-3686. https://doi.org/10.1098/rspb.2012.0705
- Ezard, T., Fujisawa, T. & Barraclough, T.G. (2009) Splits: species' limits by threshold statistics. R Package Version 1.0-19. [program]
- Folmer, O., Black, M., Hoeh, W., Lutz, R. & Vrijenhoek, R. (1994) DNA Primers for Amplification of Mitochondrial Cytochrome c Oxidase Subunit I from Diverse Metazoan Invertebrates. Molecular Marine Biology and Biotechnology, 3 (5), 294-299.
- Hall, T.A. (1999) BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symposium Series, 41, 95-98.
- Hamilton, C.A., Formanowicz, D.R. & Bond, J.E. (2011) Species Delimitation and Phylogeography of Aphonopelma hentzi (Araneae, Mygalomorphae, Theraphosidae): Cryptic Diversity in North American Tarantulas. PLoS ONE, 6, e26207. https://doi.org/10.1371/journal.pone.0026207
- Hamilton, C.A., Hendrixson, B.E., Brewer, M.S. & Bond, J.E. (2014) An evaluation of sampling effects on multiple DNA barcoding methods leads to an integrative approach for delimiting species: A case study of the North American tarantula genus Aphonopelma (Araneae, Mygalomorphae, Theraphosidae). Molecular Phylogenetics and Evolution, 71, 79-93. https://doi.org/10.1016/j.ympev.2013.11.007
- Hebert, P.D.N., Cywinska, A., Ball, S.L. & deWaard, J.R. (2003) Biological identifications through DNA barcodes. Proceedings of the Royal Society B, 270, 313-321. https://doi.org/10.1098/rspb.2002.2218
- Hebert, P.D.N., Stoeckle, M.Y., Zemlak, T.S. & Francis, C.M. (2004) Identification of Birds through DNA Barcodes. PLOS Biology, 2, e312. https://doi.org/10.1371/journal.pbio.0020312
- Hormiga, G. (2017) The discovery of the orb-weaving spider genus Pinkfloydia (Araneae, Tetragnathidae) in eastern Australia with description of a new species from New South Wales and comments on the phylogeny of Nanometinae. Zootaxa, 4311 (4), 480-490. https://doi.org/10.11646/zootaxa.4311.4.2
- Hsiao, S.T., Chuang, S.C., Chen, K.S., Ho, P.H., Wu, C.L. & Chen, C.A. (2016) DNA barcoding reveals that the common cupped oyster in Taiwan is the Portuguese oyster Crassostrea angulata (Ostreoida; Ostreidae), not C. gigas. Scientific Reports, 6, 34057. https://doi.org/10.1038/srep34057
- Hu, Y. J. (1980). On some common species of the genus Oxyopes from China. Journal of Hunan Normal University, Natural Sciences, 1980 (1), 67-75. [in Chinese]
- Hu, J.L. (1984) The Chinese spiders collected from the fields and the forests. Tianjin Press of Science and Techniques, Tianjin, 482 pp. [in Chinese]
- Hu, J.L. (2001) Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, Zhengzhou, 658 pp. [in Chinese]
- Hu, Y.J., Liu, M.X. & Li, F.J. (1985) A description of the Oxyopes sushilae Tikader, 1965 (Araneae, Oxyopidae). Journal of Hunan Normal University, Natural Sciences, 1985 (1), 28-31.
- Hu, C.C., Wu, Y.Q., Ma, L., Chen, Y.J. & Ji, X. (2019) Genetic and morphological divergence among three closely related Phrynocephalus species (Agamidae). BMC Evolutionary Biology, 19, 114. https://doi.org/10.1186/s12862-019-1443-y
- Huber, B.A. & Dimitrov, D. (2014) Slow genital and genetic but rapid non-genital and ecological differentiation in a pair of spider species (Araneae, Pholcidae). Zoologischer Anzeiger, 253, 394-403. https://doi.org/10.1016/j.jcz.2014.04.001
- Jin, Q., Hu, X.M., Han, H.L., Chen, F., Cai, W.J., Ruan, Q.Q., Liu, B., Luo, G.J., Wang, H., Liu, X., Ward, R.D., Wu, C.-S., Wilson, J.J. & Zhang, A.B. (2018) A two-step DNA barcoding approach for delimiting moth species: moths of Dongling Mountain (Beijing, China) as a case study. Scientific Reports, 8, 14256. https://doi.org/10.1038/s41598-018-32123-9
- Kimura, M. (1980) A simple method for estimating evolutionary rates of base substitutions through comparative studies of nucleotide sequences. Journal of Molecular Evolution, 16, 111-120. https://doi.org/10.1007/BF01731581
- Kim, S.T. & Lee, S.Y. (2017) Anthropoda: Arachnida: Araneae: Oecobiidae, Oxyopidae, Cybaeidae, Dictynidae, Sparassidae, Philodromidae. Spiders II. Invertebrate Fauna of Korea, 21 (42), 1-122.
- Koch, L. (1878a) s.n. In: Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Bauer & Raspe, Nurnberg, pp. 969-1044, pls. 85-91.
- Koch, L. (1878b) Japanesische Arachniden und Myriapoden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 27, 735-798, pls. 15-16.
- Kumar, S., Stecher, G. & Tamura, K. (2016) MEGA7: Molecular Evolutionary Genetics Analysis Version 7.0 for Bigger Datasets. Molecular Biology and Evolution, 33 (7), 1870-74. https://doi.org/10.1093/molbev/msw054
- Latreille, P.A. (1804) Tableau methodique des Insectes. Nouveau Dictionnaire d'Histoire Naturelle, Paris, 24, 129-295.
- Lee, C.L. (1964) Spiders of Formosa. Da-Jiang Publishing, Taichung, 84 pp. [in Chinese]
- Lo, Y.Y. & Lin, C.P. (2016) A new record and re-description of Oxyopes sushilae (Araneae, Oxyopidae) in Taiwan. Taiwan Journal of Biodiversity, 18, 137-144.
- Lohse, K. (2009) Can mtDNA Barcodes Be Used to Delimit Species? A Response to Pons et al. (2006). Systematic Biology, 58, 439-442. https://doi.org/10.1093/sysbio/syp039
- Maloney, D., Drummond, F.A. & Alford, R. (2003) Spider predation in agroecosystems: Can spiders effectively control pest populations. Technical Bulletin 190. Department of Biological Sciences, University of Maine, Orono, 32 pp.
- Masters, B.C., Fan, V. & Ross, H.A. (2011) Species Delimitation-a Geneious Plugin for the Exploration of Species Boundaries. Molecular Ecology Resources, 11 (1), 154-57. https://doi.org/10.1111/j.1755-0998.2010.02896.x
- Meyer, C.P. & Paulay, G. (2005) DNA Barcoding: Error Rates Based on Comprehensive Sampling. PLoS Biology, 3, e422. https://doi.org/10.1371/journal.pbio.0030422
- Michalko, R., Pekar, S. & Entling, M.H. (2019) An updated perspective on spiders as generalist predators in biological control. Oecologia, 189, 21-36. https://doi.org/10.1007/s00442-018-4313-1
- Mukhtar, M.K. (2013) Description of two new species of the genus Oxyopes (Araneae: Oxyopidae) from Punjab, Pakistan. Pakistan Journal of Zoology, 45 (6), 1511-1516.
- Mukhtar, M.K. (2017) Two New Species of Oxyopes (Araneae: Oxyopidae) from Punjab, Pakistan. Pakistan Journal of Zoology, 49 (1), 143-148. https://doi.org/10.17582/journal.pjz/2017.49.1.149.154
- Nadolny, A.A., Omelko, M.M., Marusik, Y.M. & Blagoev, G. (2016) A new species of spider belonging to the Pardosa lugubris - group (Araneae: Lycosidae) from Far East Asia. Zootaxa, 4072 (2), 263-281. https://doi.org/10.11646/zootaxa.4072.2.8
- Nice, C.C. & Shapiro, A.M. (1999) Molecular and morphological divergence in the butterfly genus Lycaeides (Lepidoptera: Lycaenidae) in North America: evidence of recent speciation. Journal of Evolutionary Biology, 12, 936-950. https://doi.org/10.1046/j.1420-9101.1999.00111.x
- Nyffeler, M., Dean, D.A. & Sterling, W.L. (1987) Evaluation of the importance of the striped lynx spiders, Oxyopes salticus (Araneae: Oxyopidae), as a predator in Texas cotton. Environmental Entomology, 16 (5), 1114-1123. https://doi.org/10.1093/ee/16.5.1114
- Nyffeler, M., Dean, D.A. & Sterling, W.L. (1992) Diets, feeding specialization, and predatory role of two lynx spiders, Oxyopes salticus and Peucetia viridans (Araneae: Oxyopidae), in a Texas cotton agroecosystem. Environmental Entomology, 21 (6), 1457-1465. https://doi.org/10.1093/ee/21.6.1457
- Ono, H. & Ban, M. (2009) Oxyopidae. In: Ono, H. (Ed.), The spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Tokyo, Kanagawa, pp. 249-250.
- Paik, K.Y. (1969) The Oxyopidae (Araneae) of Korea. Theses collection commemorating the 60th Birthday of Dr. In Sock Yang, 1969, 105-127.
- Papadopoulou, A., Monaghan, M.T., Barraclough, T.G. & Vogler, A.P. (2009) Sampling Error Does Not Invalidate the Yule- Coalescent Model for Species Delimitation. A Response to Lohse (2009). Systematic Biology, 58, 442-444. https://doi. org/10.1093/sysbio/syp038
- Paradis, E. & Schliep, K. (2018) ape 5.0: an environment for modern phylogenetics and evolutionary analyses in R. Bioinformatics, 35, 526-528. https://doi.org/10.1093/bioinformatics/bty633
- Pons, J., Barraclough, T.G., Gomez-Zurita, J., Cardoso, A., Duran, D.P., Hazell, S., Kamoun, S., Sumlin, W.D. & Vogler, A.P. (2006) Sequence-based species delimitation for the DNA taxonomy of undescribed insects. Systematic Biology, 55 (4), 595-609. https://doi.org/10.1080/10635150600852011
- Prevot, V., Jordaens, K., Sonet, G. & Backeljau, T. (2013) Exploring Species Level Taxonomy and Species Delimitation Methods in the Facultatively Self-Fertilizing Land Snail Genus Rumina (Gastropoda: Pulmonata). PLoS ONE, 8, e60736. https://doi.org/10.1371/journal.pone.0060736
- Puillandre, N., Lambert, A., Brouillet, S. & Achaz, G. (2012) ABGD, Automatic Barcode Gap Discovery for primary species delimitation. Molecular Ecology, 21, 1864-1877. https://doi.org/10.1111/j.1365-294X.2011.05239.x
- R Core Team (2019) R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna. Available from: https://www.R-project.org/ (accessed 14 December 2020)
- Rambaut, A. & Drummond, A. (2016) TreeAnnotator. Version 2.4.5 as part of the BEAST software package. Available from: http://www.beast2.org (accessed 14 December 2020)
- Rambaut, A., Drummond, A.J., Xie, D., Baele, G. & Suchard, M.A. (2018) Posterior Summarization in Bayesian Phylogenetics Using Tracer 1.7. Systematic Biology, 67 (5), 901-904. https://doi.org/10.1093/sysbio/syy032
- Reid, N.M. & Carstens, B.C. (2012) Phylogenetic estimation error can decrease the accuracy of species delimitation: a Bayesian implementation of the general mixed Yule-coalescent model. BMC Evolutionary Biology, 12, 196. https://doi. org/10.1186/1471-2148-12-196
- Robinson, E.A., Blagoev, G.A, Hebert, P.D.N. & Adamowicz, S.J. (2009) Prospects for using DNA barcoding to identify spiders in species-rich genera. ZooKeys, 16, 27-46. https://doi.org/10.3897/zookeys.16.239
- Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Hˆ hna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: Efficient Bayesian Phylogenetic Inference and Model Choice Across a Large Model Space. Systematic Biology, 61 (3), 539-42. https://doi.org/10.1093/sysbio/sys029
- Rosenberg, N.A. (2007) Statistical tests for taxonomic distinctiveness from observations of monophyly. Evolution, 61, 317-323. https://doi.org/10.1111/j.1558-5646.2007.00023.x
- Ross, H.A., Murugan, S. & Sibon Li, W.L. (2008) Testing the Reliability of Genetic Methods of Species Identification via Simulation. Systematic Biology, 57, 216-230. https://doi.org/10.1080/10635150802032990
- Rozas, J., Ferrer-Mata, A., Sanchez-Del Barrio, J.C., Guirao-Rico, S., Librado, P., Ramos-Onsins, S.E. & Sanchez-Gracia, A. (2017) DnaSP 6: DNA Sequence Polymorphism Analysis of Large Data Sets. Molecular Biology and Evolution, 34 (12), 3299-3302. https://doi.org/10.1093/molbev/msx248
- Saito, S. (1933) Notes on the spiders from Formosa. Transactions of the Sapporo Natural History Society, 13, 32-61.
- Santos, A.J. (2017) The jumping lynx spider Oxyopes salticus Hentz, 1845 and its Neotropical relatives (Araneae: Oxyopidae). Zootaxa, 4216 (5), 457-481. https://doi.org/10.11646/zootaxa.4216.5.3
- Satler, J.D., Carstens, B.C. & Hedin, M. (2013) Multilocus Species Delimitation in a Complex of Morphologically Conserved Trapdoor Spiders (Mygalomorphae, Antrodiaetidae, Aliatypus). Systematic Biology, 62, 805-823. https://doi.org/10.1093/sysbio/syt041
- Sherriffs, W.R. (1955) More Oriental spiders of the genus Oxyopes. Proceedings of the Zoological Society of London, 125 (2), 297-308. https://doi.org/10.1111/j.1096-3642.1955.tb00600.x
- Shivakumar, M.S. & Kumar, D. (2010) Biological Control Potential of Male and Female Oxyopes shweta (Araenae: Oxyopidae) against Polyphagous Insect Pest Spodoptera litura. Journal of Ecobiotechnology, 2 (5), 20-24.
- Song, D.X. (1991) On lynx spiders of the genus Oxyopes (Araneae: Oxyopidae) from China. Sinozoologia, 8, 169-181.
- Song, D.X., Chen, J. & Zhu, M.S. (1997) Arachnida: Araneae. In: Yang, X.K. (Ed.), Insects of the Three Gorge Reservoir area of Yangtze River. Chongqing Publishing House, Chongqing, pp. 1704-1743.
- Song, D.X., Zhu, M.S. & Li, S.Q. (1993) Arachnida: Araneae. In: Huang, C.M. (Ed.), Animals of Longqi Mountai. China Forestry Publishing House, Beijing, pp. 852-890.
- Song, D.X., Zhu, M.S. & Chen, J. (1999) The spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.
- Stamatakis, A. (2006) RAxML-VI-HPC: Maximum Likelihood-Based Phylogenetic Analyses with Thousands of Taxa and Mixed Models. Bioinformatics, 22 (21), 2688-2690. https://doi.org/10.1093/bioinformatics/btl446
- Sukumaran, J. & Knowles, L.L. (2017) Multispecies coalescent delimits structure, not species. Proceedings of the National Academy of Sciences, 114, 1607-1612. https://doi.org/10.1073/pnas.1607921114
- Swindell, S.R. & Plasterer, T.N. (1997) SeqMan: contig assembly. In: Swindell, S.R. (Ed.), Sequence data analysis guidebook. Human Press, Totowa, New Jersey, pp. 75-89. https://doi.org/10.1385/0-89603-358-9:75
- Talavera, G., Dinca, V. & Vila, R. (2013) Factors affecting species delimitations with the GMYC model: insights from a but- terfly survey. Methods in Ecology and Evolution, 4, 1101-1110. https://doi.org/10.1111/2041-210X.12107
- Tang, G. & Li, S.Q. (2012) Lynx spiders from Xishuangbanna, Yunnan, China (Araneae: Oxyopidae). Zootaxa, 3362 (1), 1-42. https://doi.org/10.11646/zootaxa.3362.1.1
- Tikader, B.K. (1965) On some new species of spiders of the family Oxyopidae from India. Proceedings of the Indian Academy of Science, 62 (3), 140-144.
- Vitt, L.J., Caldwell, J.P., Zani, P.A. & Titus, T.A. (1997) The role of habitat shift in the evolution of lizard morphology: evidence from tropical Tropidurus. Proceedings of the National Academy of Sciences, 94, 3828-3832. https://doi.org/10.1073/pnas.94.8.3828
- Welton, L.J., Travers, S.L., Siler, C.D. & Brown, R.M. (2014) Integrative Taxonomy and Phylogeny-Based Species Delimitation of Philippine Water Monitor Lizards (Varanus salvator Complex) with Descriptions of Two New Cryptic Species. Zootaxa, 3881 (3), 201-227. https://doi.org/10.11646/zootaxa.3881.3.1
- World Spider Catalog (2020) World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http://wsc. nmbe.ch/ (accessed 13 September 2020). https://doi.org/10.24436/2
- Wu, S.P., Huang, C.C., Tsai, C.L., Lin, T.E., Jhang, J.J. & Wu, S.H. (2016) Systematic revision of the Taiwanese genus Kurixalus members with a description of two new endemic species (Anura, Rhacophoridae). ZooKeys, 557, 121-153. https://doi.org/10.3897/zookeys.557.6131
- Xie, L. & Kim, J.P. (1996) Three New Species of the Genus Oxyopes from China (Araneae: Oxyopidae). Korean Arachnology, 12, 33-40.
- Xu, X., Liu, F., Chen, J., Li, D. & Kuntner, M. (2015) Integrative taxonomy of the primitively segmented spider genus Ganthela (Araneae: Mesothelae: Liphistiidae): DNA barcoding gap agrees with morphology. Zoological Journal of the Linnean Society, 175, 288-306. https://doi.org/10.1111/zoj.12280
- Yaginuma, T. (1960) Spiders of Japan in Colour. Hoikusha, Osaka, 186 pp.
- Yin, C.M., Peng, X.J., Yan, H.M., Bao, Y.H., Xu, X., Tang, G., Zhou, Q.S. & Liu, P. (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp.
- Zhang, B.S. & Zhang, F. (2018) Three new species of the spider genus Asceua from Malaysia (Araneae, Zodariidae). Zookeys, 789, 37-49. https://doi.org/10.3897/zookeys.789.24261
- Zhu, M.S. & Zhang, B.S. (2011) Spider Fauna of Henan: Arachnida: Araneae. Science Press, Beijing, 558 pp.