A revision of the species of the Neoserica (sensu lato) vulpes group (Coleoptera: Scarabaeidae: Sericini)

This paper revises the species belonging to the Neoserica (sensu lato) vulpes group and results in one new combination, Neoserica (sensu lato) vulpes (Arrow, 1946) comb. nov., and 24 new species originating mainly from south-western China: N. baishuiensis sp. nov., N. baoshana sp. nov., N. biuncinata sp. nov., N. dundai sp. nov., N. ganhaiziana sp. nov., N. heishuiana sp. nov., N. kereni sp. nov., N. laocaiana sp. nov., N. lateriuncinata sp. nov., N. leiboensis sp. nov., N. luzhouana sp. nov., N. ningyuanensis sp. nov., N. nykli sp. nov., N. parausta sp. nov., N. pseudovulpes sp. nov., N. rubellula sp. nov., N. ruzickai sp. nov., N. shinkaisiensis sp. nov., N. sichuanica sp. nov., N. usta sp. nov., N. weishanensis sp. nov., N. xiaguanensis sp. nov., N. kunmingensis sp. nov. and N. yangjiapingensis sp. nov. A key to species and illustrations of genitalia and habitus of adults are given, including distribution maps of all species. http://zoobank.org/urn:lsid:zoobank.org:pub:3FE6CC54-DB3A-4201-A6F9-332E4A397268


Introduction
Since the redefinition of the genus Neoserica Brenske, 1894 (Pope 1960;Ahrens 2003) many of the c.200 species so far grouped under Neoserica and not directly related to the type species Neoserica ursina (Brenske 1894) (i.e. Neoserica (sensu stricto) group; Ahrens 2003) are grouped preliminarily as Neoserica sensu lato (e.g. Ahrens 2004). This collective group was identified to be neither related to Neoserica sensu stricto (Ahrens 2003) nor being monophyletic (Ahrens and Vogler 2008) and all species included therein await taxonomic revision based on which hopefully their relationship and their correct classification can be subsequently established.
In the current study we investigate the taxonomy of the taxa closely related to the species Neoserica vulpes (Arrow 1946) comb. nov., described originally from Myanmar. The species of this group are characterized by an antennal club composed in males of four antennomeres (in female three) and by a more or less long ventral process of phallobase (which in a few species may be absent).
According to our present knowledge this species group is limited in distribution to the mountain regions south-east and east of the Tibetan highland. So far only one species was known for this group, N. vulpes. Here, 24 new species are discovered originating mainly from southern China. Redescription of syntype 1 Length. 8.8 mm, length of elytra: 6.4 mm, width: 4.6 mm. Body oblong, light reddish brown, antenna yellow, dorsal surface dull, nearly glabrous except a few long erect setae on elytra. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to strongly convex anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface weakly convex medially and shiny, finely and very densely irregularly punctate, distance between punctures less than their diameter,       punctures partly fused, with a few long erect setae; frontoclypeal suture very feebly incised and medially weakly angled; smooth area in front of eye short and approximately twice as wide as long; ocular canthus long and slender, impunctate, with a short single terminal seta. Frons dull, with fine and moderately dense punctures, with a few long setae in larger punctures on disc and beside eyes. Eyes large, ratio of diameter/interocular width: 0.78. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.5 times as long as remaining antennomeres combined, slightly reflexed. Mentum convexly elevated anteriorly.
Pronotum narrow, widest shortly before base, lateral margins nearly straight and subparallel, in anterior quarter weakly curved and narrowed anteriorly, anterior angles moderately produced and moderately acute, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely interrupted medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and dense punctures, on apex smooth, punctures with minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals convex and not densely punctate with punctures concentrated along striae, odd intervals with a few single, fine, erect setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed, metacoxa glabrous, with fine setae laterally, its apical margin weakly convex, posterior lateral angle blunt; each abdominal sternite with indistinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.45. Pygidium moderately and evenly convex, finely and moderately densely punctate, without smooth midline, punctures with short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin straight, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/4.0, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres partly finely punctate dorsally, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, without longitudinal impressions, with a strongly serrated ridge ventrally and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres Figure 10. Distribution of the species of the Neoserica (s.l.) vulpes group: N. baishuiensis, N. heishuiana, N. kereni, N. lateriuncinata, N. luzhouana, N. nykli, N. parausta, N. pseudovulpes, N. ruzickai, N. usta, N. vulpes and N. kunmingensis. combined and a quarter of its length longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.
Aedeagus. Figure 1A- Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and strongly convergent to moderately convex anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin Figure 11. Distribution of the species of the Neoserica (s.l.) vulpes group: N. baoshana, N. biuncinata, N. dundai, N. ganhaiziana, N. laocaiana, N. leiboensis, N. ningyuanensis, N. rubellula, N. shinkaisiensis, N. sichuanica, N. weishanensis, N. xiaguanensis and N. yangjiapingensis. distinctly sinuate medially; surface weakly convex medially and shiny, finely and densely punctate, distance between punctures less than their diameter, with a few long erect setae; frontoclypeal suture very feebly incised and medially weakly angled; smooth area in front of eye approximately twice as wide as long; ocular canthus long and slender, impunctate, with a short single terminal seta. Frons dull, with fine and moderately dense punctures, with a few long setae in larger punctures on disc and beside eyes. Eyes large, ratio of diameter/interocular width: 0.73. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.5 times as long as remaining antennomeres combined, slightly reflexed. Mentum convexly elevated anteriorly.
Pronotum narrow, widest at base, lateral margins nearly straight and subparallel, in anterior quarter weakly curved and narrowed anteriorly, anterior angles weakly produced and moderately acute, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely interrupted medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and irregularly dense punctures, on apex smooth, punctures with minute setae.
Elytra oblong, widest at middle, striae distinctly impressed, finely and densely punctate, intervals moderately convex and not densely punctate, punctures concentrated along striae, odd intervals with a few single, fine, erect setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed, metacoxa glabrous, with fine setae laterally, its apical margin weakly convex, posterior lateral angle blunt; each abdominal sternite with indistinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.62. Pygidium lost in holotype.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin straight, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.4, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres partly finely punctate dorsally, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, without longitudinal impressions, with a strongly serrated ridge ventrally and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of aedeagus very similar to N. vulpes; it differs from N. vulpes in the narrower left paramere (dorsal view), which is less blown up and less spherical (lateral view), the ventral process of phallobase is not widened apically.

Etymology
The new species is named after its type locality, Yangjiaping (Latin adjective in the nominative singular). Description Length. 7.5 mm, length of elytra: 5.5 mm, width: 3.8 mm. Body oblong, light reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal and moderately wide, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anterior margin deeply sinuate medially; surface flat and shiny, finely and densely punctate, partly punctures fusing with each other, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with one short terminal seta. Frons dull, with fine and sparse punctures, with sparse but evenly scattered long setae. Eyes moderately large, ratio of diameter/interocular width: 0.62. Antenna composed of 10 antennomeres; club with four antennomeres, 1.5 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest a quarter before base, lateral margins nearly evenly convex, weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and acute, posterior angles blunt, sharply toothlike produced at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae in punctures.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle blunt and slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.45. Pygidium weakly convex, finely and moderately densely punctate, without smooth midline, punctures with short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.1, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with three fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres, except first mesotarsomere impunctate and glabrous dorsally, with evenly spaced, short setae ventrally; metatarsomeres ventrally glabrous and with a strongly serrated ridge, with a fine longitudinal carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
Neoserica ruzickai sp. nov. differs from N. vulpes by more widely and spherically enlarged parameres, the bifid right paramere and the stouter phallobase.

Etymology
The new species is named after one of its collectors, Jan Růžička (noun in the genitive case). Labroclypeus small, subtrapezoidal, little wider than long, widest at base, lateral margins convex and strongly convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anterior margin distinctly sinuate medially; surface slightly convex medially and moderately shiny, finely, superficially and moderately densely punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, impunctate, with one short terminal seta. Frons dull, with fine and sparse punctures, with sparse and coarser punctures interspersed each bearing a long seta. Eyes moderately large, ratio of diameter/interocular width: 0.91. Antenna composed of 10 antennomeres; club with four antennomeres, 1.5 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum wide, widest at middle, lateral margins in basal half nearly subparallel, nearly evenly convex anteriorly and weakly narrowed towards moderately produced and acute anterior angles, posterior angles blunt, not rounded at the tip, anterior margin convexly produced medially, marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.56. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with minute setae, with numerous long setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, dorsally not serrated, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.2, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with three fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Mesotibia and mesotarsomeres missing in holotype, first mesotarsomere impunctate and glabrous dorsally, with evenly spaced, short setae ventrally, slightly longer than dorsal tibial spur; subsequent metatarsomeres missing in holotype. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of aedeagus rather similar to N. ruzickai sp. nov.; it differs from the latter and from all other species of the N. vulpes group by the strongly enlarged, spherical right paramere having the additional dorsal process directed distally and slightly widened and dorsoventrally flattened and rounded at apex ( Figure 2C).

Etymology.
The new species is named after its type locality, ShinKaiSi (Latin adjective in the nominative singular).

Description
Length. 9.0 mm, length of elytra: 6.2 mm, width: 4.7 mm. Body oblong, dark brown, labroclypeus, legs and margins of pronotum lighter, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus small, subtrapezoidal and moderately wide, widest at base, lateral margins nearly straight and strongly convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anterior margin distinctly sinuate medially; surface slightly convex medially and moderately shiny, finely, superficially and moderately densely punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, impunctate, with one short terminal seta. Frons dull, with fine and sparse punctures, with sparse but evenly scattered long setae. Eyes moderately large, ratio of diameter/interocular width: 0.68. Antenna composed of 10 antennomeres; club with four antennomeres, 1.3 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum wide, widest at a quarter before base, lateral margins nearly evenly convex, weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and acute, posterior angles blunt, sharply tooth-like produced at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.52. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, punctures with minute setae, with numerous long setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.8, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with three fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres, except first mesotarsomere impunctate and glabrous dorsally, with evenly spaced, short setae ventrally; first mesotarsomere with two single long setae; metatarsomeres with only very minute setae ventrally, with a strongly serrated ventral ridge and with a fine longitudinal carina immediately beside it, dorsal surface with superficial longitudinal wrinkles; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from N. ruzickai sp. nov. and N. shinkaisiensis sp. nov. (both with bifid right paramere) by having the right paramere less widened and spherical, its medial apical process strongly bent ventrally (not straight).

Etymology
The new species is named after its occurrence in the Weishan mts (Latin adjective in the nominative singular).

Description
Length. 9.4 mm, length of elytra: 6.8 mm, width: 5.1 mm. Body oblong, dark reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins straight and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface weakly convex medially and shiny, finely and densely punctate, partly punctures fusing with each other, with a few long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately twice as wide as long; ocular canthus short and triangular, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae. Eyes small, ratio of diameter/interocular width: 0.58. Antenna composed of 10 antennomeres; club with four antennomeres, as long as the remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest shortly before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and strongly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with a narrow rim of short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle nearly right-angled; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.46. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.2, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately coarse, moderately dense punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres smooth and glabrous, with evenly spaced, short setae ventrally; metatarsomeres with a strongly serrated ventral ridge and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from the previous one by the ventral process of phallobase bent laterally, the right paramere being simple and not divided into two long lobes, and the left paramere being distinctly bifid, but it is not spherically widened as in N. vulpes or N. yangjiapingensis sp. nov.; the processes of the left paramere are at least half as long as the paramere.

Etymology
The new species is named after its type locality, Ningyuan (Latin adjective in the nominative singular).

Description
Length. 9.4 mm, length of elytra: 6.4 mm, width: 5.0 mm. Body oblong, reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface weakly convex medially and shiny, finely and densely punctate, partly punctures fusing with each other, with a few long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately twice as wide as long; ocular canthus short and triangular, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes small, ratio of diameter/interocular width: 0.54. Antenna composed of 10 antennomeres; club with four antennomeres, slightly longer than remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum narrow, widest a quarter before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and strongly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border chitinous, with a very narrow rim of short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.42. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.48, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres smooth and glabrous, with evenly spaced, short setae ventrally; metatarsomeres with a strongly serrated ventral ridge and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is quite similar in shape of aedeagus to N. ningyuanensis sp. nov., it differs from N. ningyuanensis by the narrower parameres, and the ventral lobe of left paramere being not truncate as in N. ningyuanensis but simply convexly pointed.

Etymology
The new species is named after its type locality, Heishui (Latin adjective in the nominative singular). Description Length. 7.7 mm, length of elytra: 5.9 mm, width: 4.2 mm. Body oblong, dark yellowish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to weakly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface nearly flat and shiny, finely and densely, irregularly punctate, with a numerous long erect setae in coarser but superficial punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with moderately dense and long setae. Eyes moderately small, ratio of diameter/interocular width: 0.65. Antenna with 10 antennomeres; club with four antennomeres, little longer than remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins evenly curved and weakly narrowed anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae, odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.41. Pygidium moderately convex, finely and moderately densely punctate, with wide, smooth midline, with sparse, long setae on apical half. Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/4.2, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from all other species of the N. vulpes group by having the parameres distinctly shorter than the ventral process of phallobase (Figure 3).

Etymology
The new species is named after one of its collectors, Huang Keren (noun in the genitive case).
Variation Length. 7.7-8.0 mm, length of elytra: 5.9-6.1 mm, width: 4.0-4.2 mm. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes small, ratio of diameter/interocular width: 0.59. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at posterior third, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae, odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.6. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, with sparse long setae on apical half. Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.8, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The aedeagus of the new species is very similar to N. kereni, it differs from N. kereni by the strongly z-shaped and double-bent, ventral process of phallobase which in N. kereni is less strongly curved.

Etymology
The new species is named after its type locality, Baoshan (Latin adjective in the nominative singular).

Description
Length. 9.0 mm, length of elytra: 6.4 mm, width: 4.4 mm. Body oblong, reddish brown, frons darker, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to weakly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface nearly flat and shiny, finely and densely punctate, with a numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, very finely and sparsely punctate, terminal seta lacking. Frons dull, with fine and sparse punctures, with moderately dense and long erect setae beside eyes and behind frontoclypeal suture, on disc with a few shorter, semi-erect setae. Eyes large, ratio of diameter/ interocular width: 0.71. Antenna with 10 antennomeres; club with four antennomeres, 1.5 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest shortly behind middle, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and moderately sharp, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and moderately densely punctate, punctures concentrated along striae except on interval 2, odd intervals with some single, fine, semi-erect setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.7. Pygidium moderately convex, finely and densely punctate, without impunctate midline, with sparse, long setae at apex.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.7, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a fine, moderately dense punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres dorsally glabrous, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in genital shape rather similar to N. kereni, it differs from N. kereni in the less strongly bent and more sharply pointed ventral process of phallobase ( Figure 4B) as well as the distinctly longer parameres of which the left one is fused dorsally with the phallobase.

Etymology
The new species is named after its type locality Kunming (= Yunnan Fu) (Latin adjective in the nominative singular).
( Figures 4E-H Labroclypeus subrectangular, short and wide, widest at base, lateral margins straight and weakly convergent to widely rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin shallowly sinuate medially; surface weakly convex medially and shiny, finely and densely punctate, partly punctures fusing with each other, with a few long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae. Eyes moderately large, ratio of diameter/interocular width: 0.72. Antenna composed of 10 antennomeres; club with four antennomeres, 1.2 times as long as the remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest shortly before base, lateral margins evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and sharp, posterior angles blunt and strongly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with a narrow rim of short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle nearly right-angled but rounded at tip; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.37. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.94, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately coarse, moderately dense punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres smooth and glabrous, only mesotarsomeres with a very few fine, dorsal setae, with evenly spaced, short setae ventrally; metatarsomeres with a strongly serrated ventral ridge and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly shorter dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from all other so far known species of the N. vulpes group by the deeply bifurcate right paramere (median process partly hided under the left paramere) and by the ventral process of phallobase possessing a small dorsal hook at apex ( Figure 4E).

Etymology
The new species is named with reference to its brown colour with the combined Latin adjective, composed of the prefix para-(derived from Greek; near) and ustus (from Latin, tanned, browned) (Latin adjective in the nominative singular).

Description
Length. 8.1 mm, length of elytra: 6.0 mm, width: 4.8 mm. Body oblong, dark reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins slightly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin shallowly sinuate medially; surface weakly convex medially and shiny, finely and densely punctate, partly punctures fusing with each other, with a few long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside the eyes and behind frontoclypeal suture. Eyes moderately large, ratio of diameter/interocular width: 0.73. Antenna composed of 10 antennomeres; club with four antennomeres, 1.5 times as long as the remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins weakly evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and sharp, posterior angles blunt and weakly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with a narrow rim of short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle nearly right-angled; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.39. Pygidium flat, at apex strongly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.94, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately coarse, moderately dense punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres smooth and glabrous, only mesotarsomeres with a very few fine, dorsal setae, with evenly spaced, short setae ventrally; metatarsomeres with a strongly serrated ventral ridge and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly shorter dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of genitalia rather similar to N. parausta sp. nov., but the phallobase is more robust and stout with a more robust ventral process; the external process of the right paramere is distinctly shorter than the median one.

Etymology
The new species is named with reference to its body colour with the Latin adjective rubellulus (somewhat reddish) (Latin adjective in the nominative singular).
Labroclypeus subtrapezoidal, widest at base, lateral margins nearly straight and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes large, ratio of diameter/interocular width: 0.7. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest a quarter before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.5. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.76, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from all other species of the N. vulpes group by the ventral process of phallobase having at apex sharp hooks (one dorsally, one ventrally); the ventral process of phallobase is basally only weakly enlarged and not dorsoventrally produced on right side, its width at base subequal to less than one third of phallobase length.

Etymology
The new species is named after its type locality, the Ganhaizi pass (Latin adjective in the nominative singular).

Description
Length. 9.1 mm, length of elytra: 6.8 mm, width: 5.0 mm. Body oblong, reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal and moderately wide, widest at base, lateral margins nearly straight and moderately convergent to weakly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, coarsely and moderately densely, irregularly punctate, with a few long erect setae; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, without a terminal seta. Frons dull, with fine and somewhat irregularly scattered moderately dense punctures, with a few long setae beside eyes and a few short ones on disc. Eyes large, ratio of diameter/interocular width: 0.76. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and slightly rounded at tip, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures, medially narrowly smooth, minute setae present.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin slightly concave and external apical angle nearly right-angled; each abdominal sternite with indistinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.42. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.9, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres with slight superficial, longitudinal wrinkles but glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from the similar N. ganhaiziana sp. nov. by the longer ventral process of phallobase.

Etymology
The new species is named with reference to the shape of its aedeagus, with the Latin adjective (in nominative) composed of the prefix bi-(twice) and uncinatus (hookshaped) (Latin adjective in the nominative singular). Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes large, ratio of diameter/interocular width: 0.74. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest a quarter before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae. Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae, penultimate lateral interval with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.53. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae on apex.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.4, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of aedeagus similar to N. gainhaiziana sp. nov.; it differs from N. gainhaiziana in the longer and slenderer ventral process of phallobase as well as in shape of parameres: the left parameres is apically bent ventrally ( Figure 5I).

Etymology
The new species is named after one of its collectors, Stanislav Nykl (noun in the genitive case).

Variation
Length. 8.3-8.5 mm, length of elytra: 6.1-6.2 mm, width: 4.4-4.5 mm. Female: antennal club composed of three antennomeres, as long as remaining antennomeres combined. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to strongly convex anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins strongly reflexed, anterior margin deeply sinuate medially; surface strongly convex medially and shiny, finely and densely irregularly punctate, distance between punctures less than their diameter, punctures partly fused, with a few long erect setae; frontoclypeal suture very feebly incised and medially weakly angled; smooth area in front of eye short and approximately 1.5 times as wide as long; ocular canthus long and slender, impunctate, with two moderately long single terminal setae. Frons dull, with fine and dense punctures, with numerous long setae in larger punctures on disc and beside eyes. Eyes large, ratio of diameter/interocular width: 0.7. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.7 times as long as remaining antennomeres combined, strongly reflexed. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at middle, lateral margins evenly and weakly convex, in anterior third strongly narrowed anteriorly, in posterior half only weakly narrowed, anterior angles moderately produced and moderately acute, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely interrupted medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures, on apex smooth, punctures with minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate with punctures except in the second one concentrated along the striae, odd intervals with a few single, fine, erect setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed, metacoxa glabrous, with fine setae laterally, its apical margin straight, posterior lateral angle nearly right-angled; each abdominal sternite with indistinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.44. Pygidium at apex strongly convex, finely and moderately densely punctate, without smooth midline, punctures with minute setae, with numerous longer setae beside apical margin and a few on the disc. Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin straight, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.7, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with coarse and dense punctures each bearing a short adjacent seta; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres partly finely punctate dorsally, with a few long setae, with sparse, short setae ventrally; metatarsomeres dorsally impunctate, without longitudinal impressions, with a strongly serrated ridge ventrally and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from the other so far known species of the N. vulpes group by the laterally produced and carinate ventral process of phallobase which bears no hooks; parameres as in N. ganhaiziana sp. nov. more or less dorsoventrally flattened, but together they are distinctly wider ( Figure 6B).

Etymology
The new species is named according its occurrence in Lao Cai province, Vietnam (Latin adjective in the nominative singular). Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to strongly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, impunctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes. Eyes large, ratio of diameter/interocular width: 0.8. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest a quarter before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, marginal line widely missing medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior border sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest at posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, punctures concentrated along striae, penultimate lateral interval with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.48. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae at apex.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.7, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in external morphology and shape of aedeagus similar to N. ganhaiziana sp. nov. It differs from N. ganhaiziana in shape of the ventral process of phallobase which is much wider at base extending also over the left side of ventral phallobase and being strongly bent at middle ( Figure 6E, G); the left paramere is in N. pseudovulpes sp. nov. longer and more evenly curved.

Etymology
The name of the new species is composed of the latinized Greek prefix pseudo-(false) and vulpes (species name of Serica vulpes Arrow, with reference to its general similarity with this species) (noun in apposition).

Description
Length. 8.1 mm, length of elytra: 6.0 mm, width: 4.7 mm. Body oblong, reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal and moderately wide, widest at base, lateral margins weakly convex and moderately convergent to widely rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anterior margin distinctly sinuate medially; surface nearly flat and moderately shiny, finely and densely punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately nearly as wide as long; ocular canthus long and slender, impunctate, with one short terminal seta. Frons dull, with fine and sparse punctures, with sparse but evenly scattered long setae. Eyes moderately large, ratio of diameter/interocular width: 0.58. Antenna composed of 10 antennomeres; club with four antennomeres, 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum wide, widest at base, lateral margins nearly evenly convex, weakly narrowed anteriorly, anterior angles moderately produced and acute, posterior angles blunt, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border chitinous, with a very tiny rim of short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.44. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, punctures with minute setae, with numerous long setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.5, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with three fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally superficially sparsely punctate and glabrous, with evenly spaced, short setae ventrally; metatarsomeres with a strongly serrated ventral ridge and a fine longitudinal carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of aedeagus most similar to N. pseudovulpes sp. nov., it differs from N. pseudovulpes by the longer and more robustly developed ventral process of phallobase being ventrally and laterally strongly widened and deeply concave on its median side.

Etymology
The new species is named with reference to its brown colour with the Latin adjective, ustus (from Latin, tanned/browned) (Latin adjective in the nominative singular). Description Length. 8.2 mm, length of elytra: 6.4 mm, width: 4.7 mm. Body oblong, reddish brown, frons slightly darker, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins slightly convex and moderately convergent to weakly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin broadly and deeply sinuate medially; surface flat and shiny, finely and very densely punctate, partly punctures fusing with each other, with a numerous long erect setae in coarser but superficial punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, terminal seta absent. Frons dull, with fine and very sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes large, ratio of diameter/interocular width: 0.78. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.5 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest shortly before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest at posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and sparsely punctate, punctures concentrated along striae, odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.48. Pygidium moderately convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.4, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres dorsally with a few longitudinally impressed superficial punctures, glabrous, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is very similar to N. usta sp. nov. in shape of aedeagus. It differs from N. usta, however, by the ventral process of phallobase, having it dorsoventrally extended and longitudinally bent and divided into a dorsal and a ventral sublobe ( Figure 7C); the right paramere of N. sichuanica sp. nov. is distinctly longer.

Etymology
The new species is named according to its occurrence in Sichuan (Latin adjective in the nominative singular).

Description
Length. 9.8 mm, length of elytra: 7.2 mm, width: 5.1 mm. Body oblong, yellowish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins nearly straight and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes large, ratio of diameter/interocular width: 0.76. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and slightly rounded at tip, posterior angles blunt and rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.4. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.2, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres laterally slightly flattened, dorsally with slight superficial, longitudinal wrinkles but glabrous, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is in shape of aedeagus similar to that of N. sichuanica sp. nov. but its left paramere is only half as wide as long.
Description Length. 7.9 mm, length of elytra: 5.9 mm, width: 4.5 mm. Body oblong, reddish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to strongly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface weakly convex and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, impunctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, interspersed with some coarser ones bearing each a long seta, setae beside eyes slightly longer. Eyes large, ratio of diameter/interocular width: 0.77. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.7 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins subparallel in basal half, evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, marginal line widely missing medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures, impunctate along basal midline, minute setae present.
Elytra oblong, widest at posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, punctures concentrated along striae, odd intervals with some single, fine, semi-erect setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.58. Pygidium strongly but evenly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae at apex. Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.4, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally but with distinct longitudinal impressions, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs in shape of aedeagus from all other species of the N. vulpes group: the left paramere is basally strongly blown up to a subspherical appendix with a sharply pointed distal process.

Etymology
The new species is named after its occurrence in Luzhou (Latin adjective in the nominative singular).

Variation
Length. 7.9-8.0 mm, length of elytra: 5.9-6.2 mm. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to strongly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, finely and densely, irregularly punctate, with numerous long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus moderately long and slender, impunctate, with two fine terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and on disc. Eyes large, ratio of diameter/interocular width: 0.73. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins subparallel in basal half, evenly curved and weakly narrowed anteriorly, anterior angles moderately produced and sharp, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, marginal line widely missing medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders sparsely setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest at posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, punctures concentrated along striae, penultimate lateral interval with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.48. Pygidium strongly but evenly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae at apex. Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.6, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs in shape of aedeagus from all other species of the N. vulpes group: the phallobase has a long ventral beside the long dorsal process.

Etymology
The new species is named after its type locality, Xiaguan (Latin adjective in the nominative singular).
Labroclypeus shortly subtrapezoidal and wide, widest at base, lateral margins straight and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface weakly convex medially and moderately shiny, coarsely and very densely irregularly punctate, distance between punctures less than their diameter, punctures partly fused, with a few long erect setae; frontoclypeal suture very feebly incised and medially weakly angled; smooth area in front of eye short and approximately twice as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a moderately short single terminal seta. Frons dull, with fine and somewhat irregularly scattered moderately dense punctures, with a few long setae beside eyes and a few short ones on disc. Eyes moderately large, ratio of diameter/interocular width: 0.64. Antenna composed of 10 antennomeres; club with four antennomeres, nearly 1.3 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins nearly straight and weakly narrowed anteriorly, in anterior quarter weakly curved, anterior angles moderately produced and moderately acute, posterior angles blunt and not rounded at tip, anterior margin convexly produced medially, broad marginal line widely interrupted medially, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and not dense punctures, medially not smooth, minute setae present.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, odd intervals with punctures concentrated along striae and with some single fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed, metacoxa glabrous, with fine setae laterally; each abdominal sternite with indistinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.35. Pygidium strongly and evenly convex, finely and moderately densely punctate, without smooth midline, punctures with short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin straight, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/4.0, dorsal margin sharply carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with a moderately fine, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, without longitudinal impressions, with a strongly serrated ridge ventrally and a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and a quarter of its length longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new taxon can be distinguished from all other species of the N. vulpes group by the lack of the ventral process of phallobase, from all species having a phallobase without a ventral process (the following two species) by the right paramere being large and spherical, with its longitudinal extension half of phallobasal length.

Etymology
The new species is named after its type locality, Baishui (Latin adjective in the nominative singular).

Description
Length. 9.4 mm, length of elytra: 6.9 mm, width: 5.4 mm. Body oblong, light reddish brown, antenna yellow, dorsal surface with a slightly iridescent shine and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins straight and moderately convergent to distinctly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins moderately reflexed, anterior margin distinctly sinuate medially; surface very weakly convex medially and moderately shiny, finely and densely punctate, partly punctures fusing with each other, with a few long erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, without terminal seta. Frons dull, with fine and sparse punctures, with a few long setae beside eyes and behind frontoclypeal suture. Eyes moderately large, ratio of diameter/ interocular width: 0.64. Antenna composed of 10 antennomeres; club with four antennomeres, 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest at base, lateral margins nearly straight and weakly narrowed anteriorly, anteriorly weakly convex, anterior angles moderately produced and slightly rounded at tip, posterior angles blunt, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders setose; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae and odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin straight and external apical angle blunt and slightly rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/metacoxa: 1/1.53. Pygidium strongly convex, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae beside apical margin.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and moderately long, widest at apex, ratio width/length: 1/3.71, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres indistinctly punctate dorsally and glabrous, with evenly spaced, short setae ventrally; metatarsomeres ventrally glabrous and with a strongly serrated ridge, with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species is very similar to N. baishuiensis sp. nov. It differs from N. baishuiensis by the smaller right paramere lacking the interior tooth ( Figure 9B, C).

Etymology
The new species is named with reference to its lateral hook of phallobase with the combined Latin adjective, composed of the Latin noun latus-(genitive: laterisside) and uncinatus (hook-shaped) (Latin adjective in the nominative singular).

Description
Length. 8.3 mm, length of elytra: 5.6 mm, width: 4.7 mm. Body oblong, dark yellowish brown, antenna yellow, dorsal surface dull and nearly glabrous. Labroclypeus subtrapezoidal, widest at base, lateral margins weakly convex and moderately convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle, margins weakly reflexed, anterior margin distinctly sinuate medially; surface flat and shiny, coarsely and densely, irregularly punctate, with a few longer, erect setae in coarser punctures; frontoclypeal suture feebly incised and medially weakly angled; smooth area in front of eye approximately 1.5 times as wide as long; ocular canthus long and slender, very finely and sparsely punctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, with a few short setae. Eyes large, ratio of diameter/interocular width: 0.74. Antenna with 10 antennomeres; club with four antennomeres, nearly 1.2 times as long as remaining antennomeres combined. Mentum convexly elevated anteriorly.
Pronotum moderately wide, widest a quarter before base, lateral margins evenly curved and weakly narrowed anteriorly and posteriorly, anterior angles weakly produced and distinctly rounded, posterior angles blunt and slightly rounded at tip, anterior margin convexly produced medially, broad marginal line widely missing, basal margin without marginal line; surface with moderately dense and fine punctures, with minute setae only; anterior and lateral borders very sparsely setose, setae nearly completely abraded in holotype; hypomeron distinctly carinate at base. Scutellum narrow and long, sharp at apex, with fine and moderately dense punctures and minute setae.
Elytra oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals weakly convex and not densely punctate, intervals with punctures concentrated along striae, odd intervals with some single, fine setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose, apical border membranous, with short microtrichomes.
Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed; metacoxa glabrous, with fine setae laterally, apical margin convex and external apical angle rounded; each abdominal sternite with a distinct transversal row of coarse punctures each bearing a short seta between fine and moderately dense punctation. Mesosternum between mesocoxae nearly half as wide as mesofemur, with irregularly scattered, fine setae. Ratio of length of metepisternum/ metacoxa: 1/1.48. Pygidium strongly convex and shiny, finely and moderately densely punctate, without smooth midline, punctures with sparse, short setae, a few longer setae on apex.
Legs slender; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur ventrally dull, anterior margin sharply carinate, without a submarginal serrated line, posterior margin moderately convex, with a few strong setae medially, only weakly widened externally in apical half and not serrated ventrally in distal half, finely serrated dorsally, with dense, short setae. Metatibia slender and long, widest at apex, ratio width/length: 1/3.8, dorsal margin moderately carinate, with two groups of spines, basal group of spines shortly before half of metatibial length, apical one at about three quarters of metatibial length, basally with a few single fine spines; external face longitudinally convex, with moderately coarse, sparse punctures; ventral margin finely serrated, with four fine, equidistant spines; medial face impunctate, apex concavely truncate interiorly near tarsal articulation. Tarsomeres dorsally finely punctate, with a few long setae, with sparse, short setae ventrally; metatarsomeres impunctate and glabrous dorsally, with a strongly serrated ridge ventrally and glabrous, and with a fine longitudinal carina immediately beside it; first metatarsomere slightly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia long, bidentate, protarsal claws symmetrical, basal tooth of inner protarsal claw bluntly truncate apically.

Diagnosis
The new species differs from all other species of N. vulpes group by lacking the ventral process of phallobase; this feature it shares with N. lateriuncinata sp. nov. and N. baishuiensis sp. nov. which are rather similar in shape of parameres. The new species differs from these two in the shape of the dorsolateral process of the phallobase, being smaller and ventrally concavely sinuate in N. leiboensis sp. nov.

Etymology
The new species is named after its type locality close to the Leibo pass (Latin adjective in the nominative singular).
Variation Length. 7.9-8.5 mm, length of elytra: 5.5-5.7 mm, width: 4.5-4.8 mm. Female: Eyes slightly smaller, antennal club composed of four antennomeres, first joint of club subequal to one quarter of club length.