Material: Holotype, RMNH.VER.21026.a (field number RS327.1), Calcareous outcrop “Batu Batangan”, Kinabatangan River, Sabah, Borneo, approx. 5°27’35.7’’N 118°06’06.3’’E, 20 March 2012, coll. Dr. Heike Kappes, two portions covering a total of 66 slides: sagittal sections of the anterior half on 45 slides (sectioned at intervals of 7 μm); sagittal sections of the posterior half containing the copulatory apparatus on 21 slides (sectioned at intervals of 8 μm).