ENTRY       ath00906                    Pathway
NAME        Carotenoid biosynthesis - Arabidopsis thaliana (thale cress)
CLASS       Metabolism; Metabolism of terpenoids and polyketides
PATHWAY_MAP ath00906  Carotenoid biosynthesis
MODULE      ath_M00097  beta-Carotene biosynthesis, GGAP => beta-carotene [PATH:ath00906]
            ath_M00372  Abscisic acid biosynthesis, beta-carotene => abscisic acid [PATH:ath00906]
DBLINKS     GO: 0016117
ORGANISM    Arabidopsis thaliana (thale cress) [GN:ath]
GENE        AT5G17230  PSY; PHYTOENE SYNTHASE [KO:K02291] [EC:2.5.1.32]
            AT4G14210  PDS3; phytoene desaturase 3 [KO:K02293] [EC:1.3.5.5]
            AT1G10830  Z-ISO; 15-cis-zeta-carotene isomerase [KO:K15744] [EC:5.2.1.12]
            AT3G04870  ZDS; zeta-carotene desaturase [KO:K00514] [EC:1.3.5.6]
            AT1G06820  CRTISO; carotenoid isomerase [KO:K09835] [EC:5.2.1.13]
            AT5G57030  LUT2; Lycopene beta/epsilon cyclase protein [KO:K06444] [EC:5.5.1.18]
            AT3G10230  LYC; lycopene cyclase [KO:K06443] [EC:5.5.1.19]
            AT4G25700  BETA-OHASE_1; beta-hydroxylase 1 [KO:K15746] [EC:1.14.15.24]
            AT5G52570  BETA-OHASE_2; beta-carotene hydroxylase 2 [KO:K15746] [EC:1.14.15.24]
            AT1G31800  CYP97A3; cytochrome P450, family 97, subfamily A, polypeptide 3 [KO:K15747] [EC:1.14.-.-]
            AT3G53130  LUT1; Cytochrome P450 superfamily protein [KO:K09837] [EC:1.14.14.158]
            AT5G67030  ABA1; zeaxanthin epoxidase (ZEP) (ABA1) [KO:K09838] [EC:1.14.15.21]
            AT1G08550  NPQ1; non-photochemical quenching 1 [KO:K09839] [EC:1.23.5.1]
            AT1G30100  NCED5; nine-cis-epoxycarotenoid dioxygenase 5 [KO:K09840] [EC:1.13.11.51]
            AT3G14440  NCED3; nine-cis-epoxycarotenoid dioxygenase 3 [KO:K09840] [EC:1.13.11.51]
            AT3G24220  NCED6; nine-cis-epoxycarotenoid dioxygenase 6 [KO:K09840] [EC:1.13.11.51]
            AT1G78390  NCED9; nine-cis-epoxycarotenoid dioxygenase 9 [KO:K09840] [EC:1.13.11.51]
            AT4G18350  NCED2; nine-cis-epoxycarotenoid dioxygenase 2 [KO:K09840] [EC:1.13.11.51]
            AT4G19170  NCED4; nine-cis-epoxycarotenoid dioxygenase 4 [KO:K09840] [EC:1.13.11.51]
            AT1G52340  ABA2; NAD(P)-binding Rossmann-fold superfamily protein [KO:K09841] [EC:1.1.1.288]
            AT2G27150  AAO3; abscisic aldehyde oxidase 3 [KO:K09842] [EC:1.2.3.14]
            AT2G29090  CYP707A2; cytochrome P450, family 707, subfamily A, polypeptide 2 [KO:K09843] [EC:1.14.14.137]
            AT3G19270  CYP707A4; cytochrome P450, family 707, subfamily A, polypeptide 4 [KO:K09843] [EC:1.14.14.137]
            AT5G45340  CYP707A3; cytochrome P450, family 707, subfamily A, polypeptide 3 [KO:K09843] [EC:1.14.14.137]
            AT4G19230  CYP707A1; cytochrome P450, family 707, subfamily A, polypeptide 1 [KO:K09843] [EC:1.14.14.137]
            AT1G52400  BGLU18; beta glucosidase 18 [KO:K15748] [EC:3.2.1.175]
            AT1G03055  D27; beta-carotene isomerase D27-like protein [KO:K17911] [EC:5.2.1.14]
            AT2G44990  CCD7; carotenoid cleavage dioxygenase 7 [KO:K17912] [EC:1.13.11.68]
            AT4G32810  CCD8; carotenoid cleavage dioxygenase 8 [KO:K17913] [EC:1.13.11.69 1.13.11.70]
COMPOUND    C00353  Geranylgeranyl diphosphate
            C00448  trans,trans-Farnesyl diphosphate
            C02094  beta-Carotene
            C03427  Prephytoene diphosphate
            C03428  Presqualene diphosphate
            C05413  all-trans-Phytoene
            C05414  all-trans-Phytofluene
            C05421  15-cis-Phytoene
            C05430  all-trans-zeta-Carotene
            C05431  all-trans-Neurosporene
            C05432  Lycopene
            C05433  alpha-Carotene
            C05434  beta-Zeacarotene
            C05435  gamma-Carotene
            C06082  Abscisate
            C06098  Zeaxanthin
            C08579  Antheraxanthin
            C08580  Astaxanthin
            C08583  Canthaxanthin
            C08584  Capsanthin
            C08585  Capsorubin
            C08586  delta-Carotene
            C08590  Zeinoxanthin
            C08591  beta-Cryptoxanthin
            C08592  Echinenone
            C08601  Lutein
            C08606  Neoxanthin
            C08607  Neurosporaxanthin
            C08613  Torulene
            C08614  Violaxanthin
            C09190  Strigol
            C09707  Phaseic acid
            C13431  9'-cis-Neoxanthin
            C13433  9-cis-Violaxanthin
            C13453  Xanthoxin
            C13454  Xanthoxic acid
            C13455  Abscisic aldehyde
            C13456  Abscisic alcohol
            C15514  8'-Hydroxyabscisate
            C15857  9,9'-dicis-zeta-Carotene
            C15858  7,9,7',9'-tetracis-Lycopene
            C15859  1'-Hydroxy-gamma-carotene
            C15860  1'-Hydroxytorulene
            C15861  1'-Hydroxy-gamma-carotene glucoside
            C15864  1'-Hydroxy-gamma-carotene glucoside ester
            C15867  3,4-Dehydrolycopene
            C15874  3,4-Dehydrorhodopin
            C15877  Anhydrorhodovibrin
            C15878  Rhodovibrin
            C15879  Hydroxyspirilloxanthin
            C15881  Spirilloxanthin
            C15884  2-Ketospirilloxanthin
            C15885  2,2'-Diketospirilloxanthin
            C15886  3,4-Dihydroanhydrorhodovibrin
            C15887  3',4'-Dihydrorhodovibrin
            C15888  3,4,3',4'-Tetrahydrospirilloxanthin
            C15892  Chloroxanthin
            C15895  3,4-Dihydrospheroidene
            C15898  Demethylspheroidene
            C15900  Spheroidene
            C15902  Hydroxyspheroidene
            C15903  Spheroidenone
            C15905  Hydroxyspheroidenone
            C15908  Chlorobactene
            C15911  Hydroxychlorobactene
            C15914  Hydroxychlorobactene glucoside
            C15917  Hydroxychlorobactene glucoside ester
            C15935  (2'S)-Deoxymyxol 2'-(2,4-di-O-methyl-alpha-L-fucoside)
            C15937  (3R,2'S)-Myxol 2'-(2,4-di-O-methyl-alpha-L-fucoside)
            C15940  (2'S)-Deoxymyxol 2'-alpha-L-fucoside
            C15941  (3R,2'S)-Myxol 2'-alpha-L-fucoside
            C15942  (3S,2'S)-4-Ketomyxol 2'-alpha-L-fucoside
            C15943  Isorenieratene
            C15965  3'-Hydroxyechinenone
            C15966  3-Hydroxyechinenone
            C15967  Phoenicoxanthin
            C15968  Adonixanthin
            C15969  Zeaxanthin diglucoside
            C15970  Abscisic acid glucose ester
            C15971  Dihydrophaseic acid
            C15981  alpha-Cryptoxanthin
            C15982  Thermocryptoxanthin
            C15983  Thermozeaxanthin
            C15984  Thermobiszeaxanthin
            C16144  4,4'-Diapophytoene
            C16145  4,4'-Diaponeurosporene
            C16146  4,4'-Diaponeurosporenic acid
            C16147  Glucosyl-4,4'-diaponeurosporenoate
            C16148  Staphyloxanthin
            C16270  Rhodopinal
            C16271  Rhodopinal glucoside
            C16274  (2S,2'S)-Oscillol
            C16275  (2S,2'S)-Oscillol 2,2'-di(alpha-L-fucoside)
            C16276  epsilon-Carotene
            C16277  R.g.-Keto I
            C16278  R.g.-Keto III
            C16279  Thiothece-474
            C16280  Okenone
            C16282  Caloxanthin
            C16284  Nostoxanthin
            C16291  7,8-Dihydro-beta-carotene
            C16340  beta-Isorenieratene
            C16486  Astaxanthin diester
            C18036  Strigolactone ABC-rings
            C18037  5-Deoxystrigol
            C19759  7,9,9'-tricis-Neurosporene
            C19764  9,15,9'-tricis-zeta-Carotene
            C19765  15,9'-dicis-Phytofluene
            C19795  1-Hydroxy-1,2-dihydrolycopene
            C19797  4,4'-Diapolycopene
            C19798  4,4'-Diapolycopenedial
            C19892  4'-Apo-beta,psi-caroten-4'-al
            C20484  9-cis-beta-Carotene
            C20692  9-cis-10'-Apo-beta-carotenal
            C20693  Carlactone
            C21770  4,4'-Diapolycopene-4,4'-dioate
            C22058  Dihydroisopentenyldehydrorhodopin
            C22059  Isopentenyldehydrorhodopin
            C22060  Dihydrobisanhydrobacterioruberin
            C22346  1'-Hydroxy-1',2'-dihydro-beta,psi-caroten-4'-one
            C22347  Bisanhydrobacterioruberin
            C22348  Monoanhydrobacterioruberin
            C22349  Bacterioruberin
REFERENCE   
  AUTHORS   Takaichi S, Mimuro M, Tomita Y.
  TITLE     [Karotenoido-sono Tayousei to Seirikattsei] (In Japanese)
  JOURNAL   SHOKABO:109-156 (2006)
REFERENCE   
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  TITLE     [Shinpan Syokubutuhorumon no Sigunarudentatu] (In Japanese)
  JOURNAL   Syujyunsya:165-186 (2004)
REFERENCE   
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REFERENCE   PMID:9914519
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REFERENCE   PMID:15557094
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REFERENCE   PMID:12782726
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REFERENCE   PMID:10851005
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REFERENCE   PMID:12664193
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REFERENCE   PMID:7721699
  AUTHORS   Lang HP, Cogdell RJ, Takaichi S, Hunter CN.
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REFERENCE   PMID:9636041
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  TITLE     A null lesion in the rhodopin 3,4-desaturase of Rhodospirillum rubrum unmasks a cryptic branch of the carotenoid biosynthetic pathway.
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REFERENCE   PMID:7588751
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REFERENCE   PMID:16020541
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REFERENCE   PMID:8002598
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REFERENCE   PMID:10395965
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  TITLE     Functional analysis of genes from Streptomyces griseus involved in the synthesis of isorenieratene, a carotenoid with aromatic end groups, revealed a novel type of carotenoid desaturase.
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REFERENCE   PMID:11479383
  AUTHORS   Takaichi S, Maoka T, Masamoto K.
  TITLE     Myxoxanthophyll in Synechocystis sp. PCC 6803 is myxol 2'-dimethyl-fucoside, (3R,2'S)-myxol 2'-(2,4-di-O-methyl-alpha-L-fucoside), not rhamnoside.
  JOURNAL   Plant Cell Physiol 42:756-62 (2001)
            DOI:10.1093/pcp/pce098
REFERENCE   PMID:15695449
  AUTHORS   Takaichi S, Mochimaru M, Maoka T, Katoh H.
  TITLE     Myxol and 4-ketomyxol 2'-fucosides, not rhamnosides, from Anabaena sp. PCC 7120 and Nostoc punctiforme PCC 73102, and proposal for the biosynthetic pathway of carotenoids.
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REFERENCE   PMID:9092504
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  TITLE     A new type of asymmetrically acting beta-carotene ketolase is required for the synthesis of echinenone in the cyanobacterium Synechocystis sp. PCC 6803.
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REFERENCE   PMID:16242129
  AUTHORS   Mochimaru M, Masukawa H, Takaichi S.
  TITLE     The cyanobacterium Anabaena sp. PCC 7120 has two distinct beta-carotene ketolases: CrtO for echinenone and CrtW for ketomyxol synthesis.
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REFERENCE   PMID:7919981
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REFERENCE   PMID:15811329
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  TITLE     The cyanobacterium Gloeobacter violaceus PCC 7421 uses bacterial-type phytoene desaturase in carotenoid biosynthesis.
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REFERENCE   PMID:15115301
  AUTHORS   Hirabayashi H, Ishii T, Takaichi S, Inoue K, Uehara K.
  TITLE     The role of carotenoids in the photoadaptation of the brown-colored sulfur bacterium Chlorobium phaeobacteroides.
  JOURNAL   Photochem Photobiol 79:280-5 (2004)
            DOI:10.1111/j.1751-1097.2004.tb00396.x
REFERENCE   PMID:16923888
  AUTHORS   Maresca JA, Bryant DA.
  TITLE     Two genes encoding new carotenoid-modifying enzymes in the green sulfur bacterium Chlorobium tepidum.
  JOURNAL   J Bacteriol 188:6217-23 (2006)
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REFERENCE   PMID:15292122
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  TITLE     Genetic manipulation of carotenoid biosynthesis in the green sulfur bacterium Chlorobium tepidum.
  JOURNAL   J Bacteriol 186:5210-20 (2004)
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REFERENCE   PMID:9297463
  AUTHORS   Takaichi S, Wang ZY, Umetsu M, Nozawa T, Shimada K, Madigan MT.
  TITLE     New carotenoids from the thermophilic green sulfur bacterium Chlorobium tepidum: 1',2'-dihydro-gamma-carotene, 1',2'-dihydrochlorobactene, and OH-chlorobactene glucoside ester, and the carotenoid composition of different strains.
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REFERENCE   PMID:8661926
  AUTHORS   Yokoyama A, Shizuri Y, Hoshino T, Sandmann G.
  TITLE     Thermocryptoxanthins: novel intermediates in the carotenoid biosynthetic pathway of Thermus thermophilus
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REFERENCE   PMID:12763034
  AUTHORS   Hemmi H, Ikejiri S, Nakayama T, Nishino T
  TITLE     Fusion-type lycopene beta-cyclase from a thermoacidophilic archaeon Sulfolobus solfataricus.
  JOURNAL   Biochem Biophys Res Commun 305:586-91 (2003)
            DOI:10.1016/S0006-291X(03)00798-8
REL_PATHWAY ath00900  Terpenoid backbone biosynthesis
KO_PATHWAY  ko00906
///
