Chalcoderus maculatus (Fabricius, 1801), restored combination

(Fig. 1)

Ateuchus maculatus Fabricius, 1801: 64.

At. Maculatus F.” (Erichson 1847: 763) (citation).

Chalcoderus maculatus Fabricius (Erichson 1847: 763) (change of genus).

O [nthophagus] maculatus F. (Lacordaire 1855: 109 (note 3)) (citation).

Onthophagus maculatus Fabr. (Harold 1869: 1032) (citation).

O [nthophagus] maculatus Fabricius (d’Orbigny 1902: 114) (citation).

Onthophagus chloroderus d’Orbigny, 1905a: 449 (description) (new subjective synonymy) (Fig. 2)

Onthophagus chloroderus d’Orbigny (d’Orbigny 1913: 257–258) (key).

Onthophagus chloroderus d’Orbigny (d’Orbigny 1914: 657) (catalogue).

Onthophagus chloroderus d’Orb. (Boucomont & Gillet 1927: 161) (citation sensu d’Orbigny 1913).

O. maculatus F.” (Palestrini 1982: 101) (subsequent designation as type species of Chalcoderus Erichson, 1847).

Onthophagus (Parascatonomus) maculatus F. (Palestrini 1982: 98) (transfer to the subgenus Parascatonomus).

Ateuchus maculatus (Fabricius, 1801) (Branco 2011: 18) (citation as type species of Chalcoderus Erichson, 1847)

Parascatonomus (Morettoeus) chloroderus (d’Orbigny, 1905a) (Ochi & Kon 2023: 580) (transfer to Parascatonomus (Morettoeus )).

All citations of Onthophagus chloroderus published subsequent to d’Orbigny (1905a) relate to Chalcoderus maculatus (Fabricius, 1801) (see summary below in section for Chalcoderus signatipennis (de Castelnau, 1840)).

Type material examined: Ateuchus maculatus Fabricius, 1801. Lectotype (not sexed): The type series of this species comprises three specimens according to Zimsen (1984: 31). To stabilize the nomenclature, here, we designate a lectotype (Article 74), which carries the following labels “ Guinea. [Ghana] / Meier. / Mus: T. Lund / Ateuchus / maculatus. F. {handwritten on square label} // small green square // TYPE {black print on red label} // ZMUC / 00508558 // square code ZMUC Fabricius 001127 // square code WORLD SCARAB DATABASE WSD00255537 // Lectotype Ateuchus maculatus / Fabricius, 1801 Moretto, / Deschodt & Davis des. 2025 {red print} // Chalcoderus maculatus / (Fabricius, 1801) / Moretto, Deschodt & Davis / det. 2025” ZMUC. Paralectotypes (2): 1 ex. “small green square // TYPE {black print on red label} // ZMUC / 00508559 // square code ZMUCFabricius 001128 // square code WORLD SCARAB DATABASE WSD00255538 // Onthophagus / maculatus / (Fab.) / P.Moretto det. 2008 {half handwritten, half printed} // Paralectotype Ateuchus / maculatus Fabricius, 1801 / Moretto, Deschodt & Davis / det. 2025 {red print} // Chalcoderus / maculatus / (Fabricius, 1801) / Moretto, Deschodt / & Davis det. 2025”; 1 ex. “small green square // TYPE {black print on red label} // ZMUC / 00508560 // square code ZMUC Fabricius 001129 // square code WORLD SCARAB DATABASE WSD00255539 // Onthophagus / maculatus / (Fab.) / P.Moretto det. 2008 {half handwritten, half printed} // Paralectotype Ateuchus / maculatus Fabricius, 1801 / Moretto, Deschodt & Davis / det. 2025 {red print} // Chalcoderus maculatus / (Fabricius, 1801) / Moretto, Deschodt & Davis / det. 2025” (ZMUC).

Type locality: “Guinea” [Ghana] (Moretto & Génier 2015: 4)

Original description (Fabricius 1801): “maculatus. 45. A. clypeo integro, capite thoraceque cupreis, elytris testaceis nigro-maculatis. Habitat in Guinea. Mus. Dom. Lund. Statura et magnitude praecedentium. Capitis clypeus rotundatus, integer, cupreus. Thorax gibbosus, laevis, cupreus, nitidus. Elytra substriata, Testacea, strigis duabus e maculis quadratis nigris. Corpus atrum.”