(begin-cut sub4 g54)
(continue-cut g54 sub1)
(end-cut-1 sub1 sub6)
(end-cut-2 sub4 sub6)
(begin-inverse-splice sub4 sub6)
(continue-inverse-splice-1a sub1 g54)
(continue-inverse-splice-2 sub1 sub4)
(end-inverse-splice-1a g54 sub1)
(end-inverse-splice-2 g54 sub6)
(reset-1 sub4)
(begin-cut sub5 sub2)
(end-cut-1 sub2 sub3)
(end-cut-2 sub5 sub3)
(begin-transpose-splice sub3 sub4)
(end-splice-1 sub2 sub3)
(end-splice-2 sub2 sub4)
(reset-1 sub5)
(invert-single-gene-b sub1)
; cost = 4 (general cost)
