Distribution type: Native.
Distribution in Brazil: Maranhão: Centro Novo do Maranhão, Gurupi Biological Reserve, Itinga (Hernandez et al. 2018a, This study).
MARANHÃO: Centro Novo do Maranhão: LB_ ANL000277, 9 adults, in plinthosol in an old-growth forest, (3°41'11.69"S 46°45'34.87"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000278— 12 juveniles and LB_ ANL000279, 8 adults, in plinthosol in an old-growth forest, (3°41'16.35"S 46°45'34.73"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000280— 6 juveniles and LB_ ANL000281, 8 adults, in plinthosol in a logged forest, (3°40'50.95"S 46°46'13.06"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000282— 9 juveniles and LB_ ANL000283, 12 adults, in plinthosol in a logged forest, (3°40'59.74"S 46°46'1.11"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000284— 6 adults and LB_ ANL000285, 2 juveniles, in plinthosol in an old-growth forest, (3°41'27.21"S 46°45'6.19"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000286— 7 juveniles and LB_ ANL000287, 6 adults, in plinthosol in an old-growth forest, (3°41'24.87"S 46°45'13.87"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000288— 1 juvenile and LB_ ANL000289, 7 adults, in plinthosol in a logged forest, (3°41'10.36"S 46°45'56.36"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000290— 12 adults and LB_ ANL000291, 23 juveniles, in plinthosol in a logged forest, (3°41'20.33"S 46°46'1.16"W), March 2015, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000292— 1 adult and LB_ ANL000293, 1 juvenile, in plinthosol in a pasture, (3°40'25.49"S 46°47'20.47"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000294— 4 adults and LB_ ANL000295, 1 juvenile, in plinthosol in a pasture, (3°37'59.90"S 46°48'57.10"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000296, 4 adults, in plinthosol in an intermediate secondary forest, (3°41'56.61"S 46°45'57.81"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000297, 7 adults, in plinthosol in an intermediate secondary forest, (3°41'57.30"S 46°46'4.80"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000298, 9 adults, in plinthosol in an intermediate secondary forest, (3°42'7.90"S 46°45'42.63"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000299— 10 juveniles and LB_ ANL000300, 12 adults, in plinthosol in an old-growth forest,(3°41'26.11"S 46°46'14.11"W), April2016, L.M.Hernández-García, G.X.Rousseau, colls.LB_ ANL 000301— 27 juveniles, in plinthosol in an old-growth forest, (3°41'18.50"S 46°45'56.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000302— 2 juveniles and LB_ ANL000303, 11 adults, in plinthosol in an old-growth forest, (3°41'27.72"S 46°46'2.26"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000304— 29 adults and LB_ ANL000305, 61 juveniles, in plinthosol in an old-growth forest, (3°41'21.00"S 46°46'24.20"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000306— 5 adults and LB_ ANL000307, 5 juveniles, in plinthosol in an intermediate secondary forest, (3°42'18.22"S 46°45'47.40"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000308— 12 juveniles and LB_ ANL000309, 10 adults, in plinthosol in an intermediate secondary forest, (3°42'7.3"S 46°45'36.1"W), April 2016, L.M. Hernández-García, G.X. Rousseau, colls.; Itinga do Maranhão: LB_ ANL000310, 7 adults, in latosol in a pasture, (4°2'20.00"S 46°56'57.01"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000311, 1 adult, in latosol in an old-growth forest, (4°2'28.80"S 46°54'30.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000312, 2 adults, in latosol in an old-growth forest, (4°2'3.64"S 46°5'3.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000313, 3 adults, in latosol in an old-growth forest, (4°2'12.66"S 46°2'12.20"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000314— 9 juveniles and LB_ ANL000315, 1 adult, in argisol in an old-growth forest, (4°5'57.60"S 46°5'57.60"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000316— 5 adults and LB_ ANL000317, 3 juveniles, in argisol in an old-growth forest, (4°6'4.00"S 46°54'58.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000318, 2 juveniles, in argisol in a pasture, (4°4'20.98"S 46°52'59.78"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000319, 1 adult, in argisol in an intermediate secondary forest, (4°4'50.30"S 46°52'51.30"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL000320, 2 juveniles, in argisol in an intermediate secondary forest, (4°4'31.00"S 46°4'31.00"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000321— 3 juveniles and LB_ ANL000322, 7 adults, in plinthosol in a logged forest, (3°55'23.60"S 46°45'57.04"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000323— 3 adults and LB_ ANL000324, 4 juveniles, in plinthosol in a logged forest, (3°58'59.00"S 46°47'56.80"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. LB_ ANL 000325— 5 adults and LB_ ANL000326, 1 juvenile, in plinthosol in a logged forest, (3°55'3.55"S 46°45'49.55"W), April 2017, L.M. Hernández-García, G.X. Rousseau, colls. (Figure 2)