Leg 4 3-segmented with compound distal segment armed with 4 spines (I and IV (rarely III) spines, respectively); leg 1 last exopodal segment without 3 posterior/medial pinnate setae, or only single vestigial seta; leg 2 second endopodal segment with long, tapering spinules along outer margin; antenna with well-developed proximal process; small process medial to post-antennal process (Boxshall & El-Rashidy 2009; Boxshall 2018).