Published September 27, 2021 | Version v1
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An exceptional partial skeleton of a new basal raptor (Aves: Accipitridae) from the late Oligocene Namba formation, South Australia

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Mather, Ellen K., Lee, Michael S. Y., Camens, Aaron B., Worthy, Trevor H. (2021): An exceptional partial skeleton of a new basal raptor (Aves: Accipitridae) from the late Oligocene Namba formation, South Australia. Historical Biology 34 (7): 1175-1207, DOI: 10.1080/08912963.2021.1966777, URL: http://dx.doi.org/10.1080/08912963.2021.1966777

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References

  • Alley NF. 1998. Cainozoic stratigraphy, palaeoenvironments and geological evolution of the Lake Eyre Basin. Palaeogeogr Palaeoclimatol Palaeoecol. 144(3-4):239-263. doi:10.1016/S0031-0182(98)00120-5.
  • Amadon D. 1964. Taxonomic notes on birds of prey. Am Mus Novit. 2166:1-24.
  • Archer M, Tedford RH, Rich TH. 1987. The Pilkipildridae, a new family and four species of ?petauroid possums (Marsupialia: Phalangerida) from the Australian Miocene. In: Possums and opossums: studies in evolution. M Archer editor. Surrey Beatty & Sons Pty Ltd and The Royal Zoological Society of New South Wales, Sydney, Australia; p. 607-627
  • Aumann T. 2001. Habitat use, temporal activity patterns and foraging behaviour of raptors in the south-west of the Northern Territory, Australia. Wildl Res. 28:365-378. doi:10.1071/WR99091.
  • Baird RF. 1991. Avian fossils from the Quaternary of Australia. In: Vickers-Rich P, Monaghan JM, Baird RF, Rich TH, editors. Vertebrate palaeontology of Australasia. Melbourne: Pioneer Design Studio Ltd, Lilydale, & Monash University Publications Committee; p. 809-849.
  • Baird RF, Rich PV, Van Tets GF. 1991. Localities yielding avian assemblages of Quaternary age in Australia. In: Vickers-Rich P, Monaghan JM, Baird RF, Rich TH, editors. Vertebrate palaeontology of Australasia. Melbourne: Pioneer Design Studio Ltd, Lilydale, & Monash University Publications Committee; p. 850-870.
  • Barba-Montoya J, Tao Q, Kumar S. 2021. Molecular and morphological clocks for estimating evolutionary divergence times. BMC Ecol Evol, 21(1): 1-15.
  • Baumel JJ, Witmer LM (1993) Osteologia. In: Handbook of avian anatomy: nomina anatomica avium. JJ Baumel, AS King, JE Breazile, HE Evans and JC Vanden Berge editors. Publications of the Nuttall Ornithological Club 23. Nuttall Ornithological Club, Cambridge, Massachusetts; p. 45-132.
  • Beck RMD, Louys J, Brewer P, Archer M, Black KH, Tedford RH. 2020. Anew family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes). Sci Rep. 10:9741. doi:10.1038/s41598-020-66425-8
  • Boles WE. 1993. Pengana robertbolesi, a peculiar bird of prey from the Tertiary of Riversleigh, northwestern Queensland, Australia. Alcheringa. 17:19-25. doi:10.1080/03115519308619485.
  • Boles WE, Finch MA, Hofheins RH, Vickers-Rich P, Walters M, Rich TH. 2013. Afossilstone-curlew (Aves:Burhinidae) fromthe Late Oligocene/Early Miocene of South Australia. In: International meeting of the Society-for-Avian -Paleontology-and-Evolution 2012. Naturhistorisches Museum Wien, Vienna, Austria; p. 43-62.
  • Boles WE, Ivison TJ. 1999. Anew genus of dwarf megapode (Galliformes: Megapodiidae) fromthe Late Oligocene of central Australia. Smithson Cont Paleobiol. 89:199-206.
  • Brodkorb P. 1964. Catalogue of fossil birds: part 2 (Anseriformes through Galliformes). Bull Flo Mus Nat Hist. 8(3):195-335.
  • Brown L, Amadon D. eds. 1968. Eagles, hawks and falcons of the world. Michelin House, London, England.
  • Buij R. 2014. Spotted Harrier hunting lizards on foot. Aust Field Ornithol. 31 (2):107-112.
  • Bunce M, Szulkin M, Lerner HRL, Barnes I, Shapiro B, Cooper A, Holdaway RN, Penny D. 2005. Ancient DNA provides new insights into the evolutionary history of New Zealand's extinct giant eagle. PLoS Biol. 3(1):e9. doi:10.1371/ journal.pbio.0030009.
  • Burleigh JG, Kimball RT, Braun EL. 2015. Building the avian tree of life using a large-scale, sparse supermatrix. Mol Phylogenet Evol. 84:53-63. doi:10.1016/ j.ympev.2014.12.003.
  • Callen RA. 1977. Late Cainozoic environments of part of Northeastern South Australia. J Geol Soc Aust. 24(3-4):151-169. doi:10.1080/ 00167617708728976.
  • Callen RA, Tedford RH. 1976. New late Cainozoic rock units and depositional environments, Lake Frome area, South Australia. Trans R Soc S Aust. 100 (3):125-167.
  • Campbell KE, Marcus L. 1992. The relationship of hindlimb bone dimensions to body weight in birds. In: Papers in avian paleontology honoring Pierce Brodkorb. KE Campbell editor. Natural History Museum of Los Angeles County Science Series, 36, Los Angeles, California; p. 395-412
  • Campbell V, Lapointe FJ. 2009. The use and validity of composite taxa in phylogenetic analysis. Syst Biol. 58(6):560-572. doi:10.1093/sysbio/syp056.
  • Christidis L, Boles W. 2008. Systematics and taxonomy of Australian birds. Melbourne, Australia: CSIRO Publishing.
  • De Pietri V, Scofield RP, Zelenkov N, Boles WE, Worthy TH. 2016. The unexpected survival of an ancient lineage of anseriform birds into the Neogene of Australia: the youngest record of Presbyornithidae. Roy Soc Open Sci. 3:150635. doi:10.1098/rsos.150635
  • Debus SJS. 1998. The birds of prey of Australia: a field guide. Melbourne, Australia: Oxford University Press.
  • Dickinson EC, Remsen JV Jr, Eds. 2013. The Howard & Moore Complete Checklist of Birds of the World. 4th ed. Vol. 1. Eastbourne (UK): Aves Press.
  • Drexel JF, Preiss WV. 1995. The Geology of South Australia: the Phanerozoic. Vol 2. Geol. Surv. S. Aust. Bull. 54.
  • Duchene DA, Jun Tong K, Foster CSP, Duchene S, Lanfear R, Ho SYW, Kosakovsky Pond S. 2020. Linking branch lengths across sets of loci provides the highest statistical support for phylogenetic inference. Mol Biol Evol. 37(4):1202-1210. doi:10.1093/molbev/msz291.
  • Elzanowski A, Paul GS, Stidham TA. 2001. An avian quadrate from the Late Cretaceous Lance Formation of Wyoming. J Vert Paleo. 20(4):712-719. doi:10.1671/0272-4634(2000)020[0712:AAQFTL]2.0.CO;2.
  • Elzanowski A, Stidham TA. 2010. Morphology of the quadrate in the Eocene anseriform Presbyornis and extant galloanserine birds. J Morphol. 271:305-323.
  • Elzanowski A, Zelenkov NV. 2015. Aprimitive heron (Aves: Ardeidae) fromthe Miocene of Central Asia. J Ornithol. 156:837-846. doi:10.1007/s10336-015- 1164-y.
  • Eyton TC. 1867. Osteologia Avium; or, A Sketch of the Osteology of Birds. Vol 2. Hobson R, Wellington, Salop/Shropshire, England
  • Feldmann R. 1989. Whitening fossils for photographic purposes. Paleontoll Soc SpecPubl. 4:342-346. doi:10.1017/S2475262200005323.
  • Ferguson-Lees J, Christie DA. 2001. Raptors of the World. London: Christopher Helm Publishers; p. 992. eds
  • Field DJ, Lynner C, Brown C, Darroch SAF, Iwaniuk A. 2013. Skeletal correlates for body mass estimation in modern and fossil flying birds. PLoS One. 8(11): e82000. doi:10.1371/journal.pone.0082000.
  • Fordyce RE. 1983. Rhabdosteid dolphins (Mammalia: Cetacea) fromthe Middle Miocene, Lake Frome area, South Australia. Alcheringa: An Australasian Journal of Palaeontology. 7(1):27-40. doi:10.1080/ 03115518308619631.
  • Fowler DW, Freedman EA, Scannella JB, Pizzari T. 2009. Predatory functional morphology in raptors: interdigital variation in talon size is related to prey restraint and immobilisation technique. PloS One. 4:e7999. doi:10.1371/ journal.pone.0007999.
  • Gadow H. 1891. Anatomischer Theil. Dr HG Bronn's Klassen und Ordnungen des Thier-Reichs. Leipzig: wissenschaftlich dargestelit in Wort und Bild. CF Winter'sche Verlagshandlung. Vogel. Vol. 6. pt 4, p.1008.
  • Gaff P, Boles WE. 2010. Anew eagle (Aves: Accipitridae) fromthe Mid Miocene Bullock Creek Fauna of northern Australia. Rec Aust Mus. 62:71-76. doi:10.3853/j.0067-1975.62.2010.1557.
  • Gaillard C. 1939. Contribution a l'etude des oiseaux fossils. Arch Mus Hist Nat Lyon. 15:1-100.
  • Gill, B. J., Bell, B. D., Chambers, G. K., Medway, D. G., Palma, R. L., Scofield, R. P., Tennyson, A. J. D. & Worthy, T. H. (2010) Checklist of the Birds of New Zealand, Norfolk and Macquarie Islands, and the Ross Dependency, Antarctica. B. J. Bell (Ed). Te Papa Press, Wellington
  • Gill F, Donsker D, Rasmussen P (Eds). 2020. IOC World Bird List (v10.2). doi: 10.14344/IOC.ML.10.2.
  • Goloboff PA, Pittman M, Pol D, Xu X. 2019. Morphological datasets fit a common mechanism much more poorly than DNA sequences and call into question the Mkv model. Syst Biol. 68(3):494-504.
  • Griffiths CS, Barrowclough GF, Groth JG, Mertz LA. 2007. Phylogeny, diversity, and classification of the Accipitridae based on DNA sequences of the RAG-1 exon. J Avian Biol. 38:587-602. doi:10.1111/j.0908-8857.2007.03971.x.
  • Hackett SJ, Kimball RT, Reddy S, Bowie RC, Braun EL, Braun MJ, Chojnowski JL, Cox WA, Han K-L, Harshman J, et al. 2008. Aphylogenomic study of birds reveals their evolutionary history. Science. 320:1763-1768. doi:10.1126/science.1157704.
  • Holdaway RN 1991. Systematics and palaeobiology of Haast's eagle (Harpagornis moorei Haast, 1872) (Aves: Accipitridae). Unpublished PhD thesis. Department of Zoology, University of Canterbury, Christchurch (New Zealand). 472 p.
  • Holdaway RN. 1994. An exploratory phylogenetic analysis of the genera of the Accipitridae, with notes on the biogeography of the family. In: Meyburn B-U, Chancellor RD, editors. Raptor conservation today. London (UK): World Working Group on Birds of Prey and Owls; p. 601-649.
  • Hou L. 1984. The Aragonian vertebrate fauna of Xiacaswan, Jiangsu 2. Aegypinae (Falcaniformes [sic], Aves). Vert PalAs. 22(1):14-22.
  • Hou L, Zhou Z, Zhang F, Li J. 2000. Anew vulture fromthe Miocene of Shandong, eastern China. Vert PalAs. 38(2):108-112.
  • Jollie M. 1976. Acontribution to the morphology and phylogeny of the Falconiformes. Evolutionary Theory, 1: 285-298.
  • Knapp M, Thomas JE, Haile J, Prost S, Ho SYW, Dussex N, Cameron-Christie S, Kardailsky O, Barnett R, Bunce M, et al. 2019. Mitogenomic evidence of close relationships between New Zealand's extinct giant raptors and small-sized Australian sister-taxa. Mol Phylogenet Evol. 134:122-128. doi:10.1016/j. ympev.2019.01.026.
  • Kurochkin EN. 1968. Fossil remains of birds from Mongolia. Ornitologiya. 9:323-330.
  • Kurochkin EN. 1976. Asurvey of the Paleogene birds of Asia. Smithson Cont Paleobiol. 27:75-86.
  • Lanfear R, Frandsen PB, Wright AM, Senfeld T, Calcott B. 2016. PartitionFinder 2: new methods for selecting partitioned models of evolution for molecular and morphological phylogenetic analyses. Mol Biol Evol. msw260. doi:10.1093/molbev/msw260
  • Lerner HRL, Mindell DP. 2005. Phylogeny of eagles, Old World vultures, and other Accipitridae based on nuclear and mitochondrial DNA. Mol Phylogenet Evol. 37:327-346. doi:10.1016/j.ympev.2005.04.010.
  • Li Z, Clarke JA, Zhou Z, Deng T. 2016. Anew Old World vulture from the late Miocene of China sheds light on Neogene shifts in the past diversity and distribution of the Gypaetinae. Auk. 133:615-625. doi:10.1642/AUK-15-240.1.
  • Lindsay JM. 1987. Age and habitat of a monospecific foraminiferal fauna from near-type Etadunna Formation, Lake Palankarinna, Lake Eyre Basin. South Aust Department Mines Rep. 87:93.
  • Linnaeus C. 1758. Systema Naturae per Regna Tria Naturae, 10th Edition, revised, Vol 1: Regnum Animale. Salvii, L. Holmiae, Stockholm, Sweden, iv + 824 pp.
  • Livezey BC, Zusi RL. 2007. Higher-order phylogeny of modern birds (Theropoda, Aves: neornithines) based on comparative anatomy. II. Analysis and discussion. Zool J Linn Soc-Lond. 149:1-95.
  • Louys J, Price GJ. 2015. The Chinchilla Local Fauna: an exceptionally rich and well-preserved Pliocene vertebrate assemblage from fluviatile deposits of south-eastern Queensland, Australia. Acta Palaeontologica Polonica. 60 (3):551-572.
  • Marchant S, Higgins PJ. 1993. Handbook of Australian, New Zealand and Antarctic Birds. Vol. 2: Raptors to Lapwings. Melbourne (Victoria): Oxford University Press.
  • Martin HA. 1990. The palynology of the Namba Formation in the Wooltana-1 bore, Callabonna Basin (Lake Frome), South Australia, and its relevance to Miocene grasslands in central Australia. Alcheringa. 14(3):247-255. doi:10.1080/03115519008619058.
  • Mayr G. 2006a. The post-cranial osteology and phylogenetic position of the Middle Eocene Messelastur gratulator Peters, 1994 - a morphological link between owls (Strigiformes) and falconiform birds? J Vert Palaeontol. 25 (3):635-645. doi:10.1671/0272-4634(2005)025[0635:TPOAPP]2.0.CO;2.
  • Mayr G. 2006b. Anew raptorial bird from the Middle Eocene of Messel, Germany. Hist Biol. 18(2):99-106. doi:10.1080/08912960600640762.
  • Mayr G. 2009. Awell-preserved skull of the "falconiform" bird Masillaraptor from the middle Eocene of Messel (Germany). Palaeodiversity. 2:315-320.
  • Mayr G. 2011. Well-preserved new skeleton of the Middle Eocene Messelastur substantiates sister group relationship between Messelasturidae and Halcyornithidae (Aves, ?Pan-Psittaciformes). J Syst Palaeontol. 9:159-171. doi:10.1080/14772019.2010.505252.
  • Mayr G. 2014. Comparative morphology of the radial carpal bone of neornithine birds and the phylogenetic significance of character variation. Zoomorphology. 133:425-434. doi:10.1007/s00435-014-0236-5.
  • Mayr G. 2016. Variations in the hypotarsus morphology of birds, and their evolutionary significance. Acta Zool. 97(2):196-210. doi:10.1111/azo.12117.
  • Mayr G. 2017. Avian Evolution. Chichester (West Sussex): John Wiley and Sons.
  • Mayr G. 2018. Size and number of the hypoglossal nerve foramina in the avian skull and their potential neuroanatomical significance. J Morphol. 279 (2):274-285. doi:10.1002/jmor.20770.
  • Mayr G, Hurum JH. 2020. A tiny, long-legged raptor from the early Oligocene of Poland may be the earliest bird-eating diurnal bird of prey. Sci Nat. 107(48). doi:10.1007/s00114-020-01703-z.
  • Mayr G, Perner T. 2020. Anew species of diurnal birds of prey from the late Eocene of Wyoming (USA) - one of the earliest New World records of the Accipitridae (hawks, eagles and allies). Neues Jahrb Geol Palaontol-Abh. 297 (2):205-215. doi:10.1127/njgpa/2020/0921.
  • Megirian D, Prideaux GJ, Murray PF, Smit N. 2010. An Australian land mammal biochronological scheme. Paleobiology. 36(4):658-671. doi:10.1666/09047.1.
  • Migotto R (2013) Phylogeny of Accipitridae (Aves: Accipitriformes) based on osteological characters. PhD Dissertation, Universidade de Sao Paulo, Insituto de Biociencias, Departmento de Zoologia.
  • Miller AH. 1966. The fossil pelicans of Australia. Mem Qld Mus. 14(5):181-190.
  • Miller MA, Pfeiffer W, Schwartz T 2010. Creating the CIPRES Science Gateway for inference of large phylogenetic trees. In: Proceedings of the Gateway Computing Environments Workshop (GCE), New Orleans, LA; p. 1-8.
  • Milne-Edwards A. 1863. Memoire sur la distribution geologique des oiseaux fossils et description de quelques especes nouvelles. Ann Sci Nat Zool. 4 (20):133-176.
  • Milne-Edwards A. 1871. Des caracteres osteologiques des oiseaux de proie diurnes. In: Recherches anatomiques et paleontologiques pour servir a l'histoire des oiseaux fossils de la France. Victor Masson et Fils, Paris, France; p. 406-473.
  • Milne-Edwards A. 1892. Sur les oiseaux fossiles des depots Eocenes de Phosphate de Chaux du sud de la France. Comptes Rendus duSecond Congres Ornithologique International, Budapest. p. 60-80.
  • Mindell DP, Fuchs J, Johnson JA. 2018. Phylogeny, taxonomy and geographic diversity of diurnal raptors: Falconiformes, Accipitriformes and Cathartiformes. In: Sarasola JH, Grande J, Negro J, editors. Birds of Prey: biology and conservation in the XXI century. Springer, Cham, Switzerland; p. 3-32.
  • Mlikovsky J. 2002. Cenozoic birds of the world part 1: Europe. Ninox Press, Praha, Czech Republic.
  • Mourer-Chauvire C. 1991. The Horusornithidae nov. fam., Accipitriformes (Aves) with a hyperflexible intertarsal joint from the Eocene of Quercy. Geobios. 13:183-192. doi:10.1016/S0016-6995(66)80023-2.
  • Nagy J, Tokolyi J. 2014. Phylogeny, historical biogeography and the evolution of migration in accipitrid birds of prey (Aves: Accipitriformes). Ornis Hungarica. 22:15-35. doi:10.2478/orhu-2014-0008.
  • Norrish K, Pickering JG. 1983. Clay Minerals. In: Soils: an Australianviewpoint. CSIRO Division of Soils, editors. Melbourne: CSIRO/Academic Press; p. 281-308.
  • Oatley G, Simmons RE, Fuchs J. 2015. Amolecular phylogeny of the harriers (Circus, Accipitridae) indicate the role of long distance dispersal and migration in diversification. Mol Phylogenet Evol. 85:150-160. doi:10.1016/j. ympev.2015.01.013.
  • Olson SL. 1985. The fossil record of birds. In: Farmer DS, King JR, Parkes KC, editors. Avian Biology (Vol. 8). New York: Academic Press; p. 79-252.
  • Peters JL. 1934. Check-list of the birds of the world: vol. 1. Cambridge: Harvard University Press.
  • Pledge NS. 2003. Anew species of Muramura Pledge (Wynyardiidae: Marsupialia) from the Middle Tertiary of the Callabonna Basin, Northeastern South Australia. Bull Am Mus Nat Hist. 279:541-555. doi:10.1206/0003-0090(2003)279<0541: C>2.0.CO;2.
  • Pledge NS. 2016. New specimens of ektopodontids (Marsupialia: Ektopodontidae) from South Australia. Mem Mus Vic. 74:173-187. doi:10.24199/j.mmv.2016.74.15.
  • Prum RO, Berv JS, Dornburg A, Field DJ, Townsend JP, Lemmon EM, Lemmon AR. 2015. Acomprehensive phylogeny of birds (Aves) using targeted next-generation DNA sequencing. Nature. 526(7574):569-573. doi:10.1038/nature15697.
  • Rasmussen DT, Olsen SL, Simons EL. 1987. Fossil birds from the Oligocene Jebel Qatrani Formation, Fayum Province, Egypt. Smithson Cont Paleobiol. 62:1-19.
  • Rich P, van Tets J. 1982. Fossil birds of Australia and New Guinea: their biogeographic, phylogenetic and biostratigraphic input. In: The Fossil Vertebrate Record of Australasia. PV Rich and EM Thompson editors. Monash University Offset Printing Unit, Clayton, Victoria; p. 235-384.
  • Rich TH, Archer M, Hand SJ, Godthelp H, Muirhead J, Pledge NS, Flannery TF, Woodburne MO, Case JA, Tedford RH, et al. 1991. Australian Mesozoic and Tertiary terrestrial mammal localities. In: Vickers-Rich P, Monaghan JM, Baird RF, Rich TH, editors. Vertebrate palaeontology of Australasia. Melbourne: Pioneer Design Studio Ltd, Lilydale, & Monash University Publications Committee; p. 1005-1057.
  • Rich TH, Archer M, Tedford RH. 1978. Raemeotherium yatkolai, gen. et. sp. nov., a primitive diprotodontid from the medial Miocene of South Australia. Mem Mus Vic. 39:85-91. doi:10.24199/j.mmv.1978.39.06.
  • Rich THV, Archer M. 1979. Namilamadeta snideri, a new diprotodontan (Marsupialia, Vombatoidea) from the medial Miocene of South Australia. Alcheringa. 3(3):197-208. doi:10.1080/03115517908527793.
  • Ridgway R. 1874. Catalogue of the ornithological collection of the Boston Society of Natural History Part II: Falconidae. Proc Boston Soc Nat. 16:43-72.
  • Ronquist F, Klopfstein S, Vilhelmsen L, Schulmeister S, Murray DL, Rasnitsyn AP. 2012. Atotal-evidence approach to dating with fossils, applied to the early radiation of the Hymenoptera. Syst Biol. 61(6):973-999. doi:10.1093/sysbio/sys058.
  • Sharpe RB. 1874. Catalogue of the Accipitres, or diurnal birds of prey, in the collection of the British Museum (Volume 1). UK (London): Order of the Trustees.
  • Sibley CG, Ahlquist JE. 1990. Phylogeny and classification of birds: a study in molecular evolution. New Haven, Connecticut: Yale University Press.
  • Stirton RA, Tedford RH, Miller AH. 1961. Cenozoic stratigraphy and vertebrate palaeontology of the Tirari Desert, South Australia. Rec South Aust Mus. 14:19-61.
  • Stresemann E, Amadon D. 1979. Order Falconiformes. In: Mayr E, Cottrell G, editors. Check-list of the birds of the world, Volume I, 2nd edition; revision of the work of James L. Peters. Museum of Comparative Zoology, Cambridge, Massachusets. p. 271-425.
  • Sushkin PP. 1905. Zur Morphologie des Vogelskelets: vergleichende Osteologie der normalen Tagraubvogel (Accipitres) und die Fragen der Classification: mit IV Taf. Nouv MemSoc Imp Naturalistes Moscou. 16:164-247.
  • Tedford RH, Woodburne MO. 1987. The Ilariidae, a new family of vombatiform marsupials from Miocene strata of South Australia and an evaluation of the homology of molar cusps in the Diprotodontia. In: Archer M, editor. Possums and opossums: studies in evolution (Vol. 2). New South Wales: Surrey Beatty and Sons, Chipping Norton; p. 401-418.
  • Tedford RH, Archer M, Bartholomai A, Plane M, Pledge NS, Rich T, Rich P, Wells RT. 1977. The discovery of Miocene vertebrates, Lake Frome area, South Australia. BMR J Aust Geol Geop. 2:53-57.
  • Thorn KM, Hutchinson MN, Lee MSY, Brown NJ, Camens AB, Worthy TH. 2021. Anew species of Proegernia from the Namba Formation in South Australia and the early evolution and environment of Australian egerniine skinks. Roy Soc Open Sci. 8:201686. doi:10.1098/ rsos.201686.
  • Travouillon KJ, Archer M, Hand SJ, Godthelp H. 2006. Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, north-western Queensland. Alcheringa: an Australasian journal of Palaeontology,30(S1): 323-349.
  • Vickers-Rich P. 1991. The Mesozoic and Tertiary history of birds on the Australian plate. In: Vickers-Rich P, Monaghan JM, Baird RF, Rich TH, editors. Vertebrate palaeontology of Australasia. Melbourne: Pioneer Design Studio Ltd, Lilydale, & Monash University Publications Committee; p. 721-809.
  • Vieillot LJP. 1816. Analyse d'une nouvelle ornithology elementaire. D'eterville, Paris, 70 pp.
  • Vigors NA. 1824. Sketches in ornithology; or, observations on the leading affinities of some of the more extensive groups of birds. On the groups of Falconidae. Zoological Journal, 308-346
  • Walker CA, Dyke GJ. 2006. New records of fossil birds of prey from the Miocene of Kenya. Hist Biol. 18(2):95-98. doi:10.1080/08912960600639434.
  • Wetmore A. 1933. An Oligocene eagle from Wyoming. Smith Misc Collect. 87 (19):1-9.
  • Whitacre D, Jenny JP. 2013. Neotropical birds of prey: biology and ecology of a forest raptor community. Ithaca: Cornell University Press, Sage House.
  • Willis PMA. 1997. Review of fossil crocodiles from Australasia. Aust Zool. 30 (3):287-298. doi:10.7882/AZ.1997.004.
  • Willis PMA, Molnar RE. 1991. Anew middle Tertiary crocodile from Lake Palankarinna, South Australia. Rec South Aust Mus. 25(1):39-55.
  • Wink M. 1995. Phylogeny of Old and New World vultures (Aves: Accipitridae and Cathartidae) inferred from nucleotide sequences of the mitochondrial cytochrome b gene. Z Naturforsch C. 50:868-882. doi:10.1515/znc-1995-11-1220.
  • Wink M, Sauer-Gurth H. 2004. Phylogenetic relationships in diurnal raptors based on nucleotide sequences of mitochondrial and nuclear marker genes. In: Chancellor RD, Meyburg B-U, editors. Raptors Worldwide. Budapest: World Working Group on Birds of Prey and Owls, MME/Birdlife Hungary; p. 483-498.
  • Woodburne MO, Tedford RH, Archer M, Pledge NS. 1987a. Madakoala, anew genus and two species of Miocene koalas (Marsupialia: Phascolarctidae) from South Australia and a new species of Perikoala. In: Archer M, editor. Possums and opossums: studies in evolution (Vol. 1). New South Wales: Surrey Beatty and Sons, Chipping Norton; p. 293-317.
  • Woodburne MO, Tedford RH, Archer M. 1987b. New Miocene pseudocheirids (Pseudocheiridae: Marsupalia) from South Australia. In: Archer M, editor. Possums and opossums: studies in evolution (Vol. 2). New South Wales: Surrey Beatty and Sons, Chipping Norton; p. 639-679.
  • Woodburne MO, MacFadden BJ, Case JA, Springer MS, Pledge NS, Power JD, Woodburne JM, Springer KB. 1994. Land mammal biostratigraphy and magnetostratigraphy of the Etadunna Formation (Late Oligocene) of South Australia. J Vert Paleontol. 13(4):483-515. doi:10.1080/ 02724634.1994.10011527.
  • Woodburne MO, Tedford RH. 1975. The first Tertiary monotreme from Australia. Am Mus Novit. 2588:1-11.
  • Woodburne MO, Tedford RH, Archer M, Turnbull WD, Plane MD, Lundelius EL. 1985. Biochronology of the continental mammal record of Australia and New Guinea. Spec Pub South Aus Department Mines Energy. 5:347-363.
  • Woodhead J, Hand SJ, Archer M, Graham I, Sniderman K, Arena DA, Black KH, Godthelp H, Creaser P, Price E. 2016. Developing a radiometrically-dated chronologic sequence for Neogene biotic change in Australia, from the Riversleigh World Heritage Area of Queensland. Gondwana Res. 29 (1):153-167. doi:10.1016/j.gr.2014.10.004.
  • Worthy TH. 2009. Descriptions and phylogenetic relationships of two new genera and four new species of Oligo-Miocene waterfowl (Aves: Anatidae) from Australia. Zool J Linn Soc-Lond. 156:411-454. doi:10.1111/j.1096- 3642.2008.00483.x.
  • Worthy TH. 2011. Descriptions and phylogenetic relationships of a new genus and two species of Oligo-Miocene cormorants (Aves: Phalacrocoracidae) from Australia. Zool J Linn Soc-Lond. 163(1):277-314. doi:10.1111/j.1096- 3642.2011.00693.x.
  • Worthy TH, Holdaway RN. 2002. The lost world of the moa: prehistoric life of New Zealand. Bloomington, Indiana: Indiana University Press.
  • Worthy TH, Mitri M, Handley WD, Lee MS, Anderson A, Sand C, Evans AR. 2016. Osteology supports a stem-gallifo rm affinity for the giant extinct flightless bird Sylviornis neocaledoniae (Sylviornithidae, Galloanseres). PLOS ONE. 11(3):e0150871. doi:10.1371/journal.pone.0150871.
  • Worthy TH, Nguyen JMT. 2020. An annotated checklist of the fossil birds of Australia. Trans R Soc S Aust. 144(1):66-108.
  • Worthy TH, Yates A. 2018. Areview of the smaller birds from the late Miocene Alcoota local faunas of Australia with a description of a new anatids species. Contribuciones del MACN. 7:221-252.
  • Zhang Z, Feduccia A, James HF, Iwaniuk A. 2012. A Late Miocene accipitrid (Aves: Accipitriformes) from Nebraska and its implications for the divergence of Old World vultures. PLOS ONE. 7(11):e48842. doi:10.1371/journal. pone.0048842.
  • Zhang Z, Zheng X, Zheng G, Hou L. 2010. Anew Old World vulture (Falconiformes: Accipitridae) from the Miocene of Gansu Province, northwest China. J Ornithol. 151:401-408. doi:10.1007/s10336-009- 0468-1.