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Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005
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Prendini, Lorenzo, Loria, Stephanie F. (2020): Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005. Bulletin of the American Museum of Natural History 2020 (442): 1-480, DOI: 10.1206/0003-0090.442.1.1, URL: https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-442/issue-1/0003-0090.442.1.1/Systematic-Revision-of-the-Asian-Forest-Scorpions-Heterometrinae-Simon-1879/10.1206/0003-0090.442.1.1.full
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References
- Adams, A., E. Marais, S. Turner, L. Prendini, and B. Pinshow. 2016. Similar burrow architecture of three arid-zone scorpion species implies similar ecological function. Natuurwissenschaften (Science of Nature) 103: 56 (11 pp.).
- Alexander, A.J. 1958. On the stridulation of scorpions. Behaviour 12: 339-352.
- Alexander, A.J. 1959. A note on the evolution of stridulation within the family Scorpionidae. Proceedings of the Zoological Society of London 133: 391-399.
- Alexander, A.J., and D.W. Ewer. 1957. A chemo-receptor in the scorpion Opisthophthalmus. South African Journal of Science 53: 421-422.
- Amr, Z., O.A. Abed, T. Al Share, N. Hamidan, and L. Prendini. 2015. New records of Jordanian scorpions. Jordan Journal of Natural History 2: 30-38.
- Amr, Z.S., and R. El-Oran. 1994. Systematics and distribution of scorpions (Arachnida, Scorpionida) in Jordan. Bolletino di Zoologia 61: 185-190.
- Aswathi, K., and P.M. Sureshan. 2017. Additions to the scorpion fauna (Arachnida: Scorpiones) of Kerala, India, with an illustrated key to the genera. Journal of Threatened Taxa 9: 9844-9850.
- Auber, M. 1963. Reproduction et croissance de Buthus occitanus Amx. Annales des Sciences Naturelles, Zoologie, Paris 5: 273-286.
- Ausserer, A. 1880. Arachnida. Zoologischer Jahresbericht 1879: 430-470.
- Axelrod, D.I., and P.H. Raven. 1978. Late Cretaceous and Tertiary vegetation history of Africa. In M.J.A. Werger (editor), Biogeography and ecology of southern Africa 1: 77-130. The Hague: W. Junk.
- Banks, N. 1928. Scorpiones and Pedipalpi collected by Dr. E. Mjoberg in Borneo. Journal of Sarawak Museum 3: 505-506.
- Bardi, J.K., and C.J. George. 1943. Digestive glands of the scorpion. A physiological study. Journal of the University of Bombay n.s. B 11: 91-115.
- Bastawade, D.B. 1992. Morphological study of spermatophore of a common Indian scorpion Mesobuthus tamulus tamulus (Fabr.). Records of the Zoological Survey of India 91: 221-225.
- Bastawade, D.B. 2002. Scorpion diversity. In A.P Jagtap and N.P. Singh (editors), Biodiversity of the Western Ghats of Maharashtra. Current Knowledge: 94, 95, 294-296. Dehra Dun: Bishen Singh Mahendra Pal Singh.
- Bastawade, D.B. 2004. Scorpionida, Araneae and Solifugae. In Fauna of Pench National Park. Zoological Survey of India, Conservation Area Series 20: 285- 312.
- Bastawade, D.B. 2005. Arachnida: Scorpionida and Solifugae. In Fauna of Melghat Tiger Reserve. Zoological Survey of India, Conservation Area Series 24: 411-419.
- Bastawade, D.B. 2006a. Arachnida: Scorpionida and Amblypygi. In Fauna of Sanjay Gandhi National Park (Invertebrates). Zoological Survey of India, Conservation Area Series 26: 131-138.
- Bastawade, D.B. 2006b. Arachnida: Scorpionida and Solifugi. In Fauna of Tadoba Andhari Tiger Reserve. Zoological Survey of India, Conservation Area Series 25: 283-309.
- Bastawade, D.B. 2008. Arachnida: Scorpionida, Araneae and Opiliones. In Fauna of Lonar Wildlife Sanctuary. Zoological Survey of India, Conservation Area Series 37: 133-153.
- Bastawade, D.B. 2009. Arachnida: Scorpiones. In Fauna of Bhimashankar Wildlife Sanctuary. Zoological Survey of India, Conservation Area Series 42: 215-222.
- Bastawade, D.B., and M. Borkar. 2008. Arachnida (Orders Scorpiones, Uropygi, Amblypygi, Araneae and Phalangida). In Fauna of Goa. Zoological Survey of India, State Fauna Series 16: 211-242.
- Bastawade, D.B., P.M. Sureshan, and C. Radhakrishnan. 2004. An illustrated key to the identification of scorpions (Scorpionida: Arachnida) of Kerala and notes on some interesting new records. Records of the Zoological Survey of India 103: 43-58.
- Bastawade, D.B., P.M. Sureshan, and C. Radhakrishnan. 2005. A new subfamily, genus and species of scorpion (Arachnida: Scorpionida) from India. Records of the Zoological Survey of India 104: 77-82.
- Becker, L. 1880. Etudes sur les scorpions. Annales de la Societe Entomologique de Belgique 25: 134-145.
- Bellevoye, A. 1870. Dons divers faits au Cabinet d'Histoire Naturelle de Metz et description des especes nouvelles qu'ils contiennent. Bulletin de la Societe d'Histoire Naturelle du Department de la Moselle (Metz) 12: 105-114.
- Birula, A.A. 1898. Ein Beitrag zur Kenntniss der Skorpionenfauna Kleinasiens. Horae Societatis Entomologicae Rossicae 33: 132-140. [preprint: October 1898; collated issue: 1901]
- Birula, A.A. 1910. Uber Scorpio maurus Linne und seine Unterarten. Horae Societatis Entomologicae Rossicae 35: 115-192.
- Birula, A.A. 1917a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya [Memoires du Musee du Caucase]. Tiflis: Imprimerie de la Chancellerie du Comite pour la Transcaucasie A (5): 1-253. [in Russian; English translation: A.A. Byalynitskii-Birulya. 1964. Arthrogastric arachnids of Caucasia. 1. Scorpions. Jerusalem: Israel Program for Scientific Translations, 170 pp.]
- Birula, A.A. 1917b. Faune de la Russie et des pays limitrophes fondee principalement sur les collections du Musee Zoologique de l'Academie des Sciences de Russie. Arachnides (Arachnoidea). Petrograd 1 (1): xx, 1-227. [in Russian; English translation: A.A. Byalynitskii-Birulya. 1965. Fauna of Russia and Adjacent Countries. Arachnoidea. Vol. I. Scorpions. Jerusalem: Israel Program for Scientific Translations, xix, 154 pp.]
- Birula, A.A. 1925. Skorpiologische Beitrage. 10. Zur geographischen Verbreitung zweier weitverbreiteter Skorpionen-Arten Palaearcticums. Zoologischer Anzeiger 63: 93-96.
- Biswas, A.T. 1984. Scorpions (Scorpiones). In Fauna of Rajaji National Park. Zoological Survey of India, Fauna of Conservation Areas 5: 317-323.
- Booncham, U., D. Sitthicharoenchai, A.O. Pradatsundarasar, S. Prasarnpun, and K. Thirakhupt. 2007. Sexual dimorphism in the Asian giant forest scorpion, Heterometrus laoticus Couzijn, 1981. NU International Journal of Science 4: 42-52.
- Borelli, A. 1915. Gli scorpioni del Museo Civico di Storia Naturale di Milano. Atti della Societa Italiana di Scienze Naturali 53: 456-464.
- Braunwalder, M.E., and V. Fet. 1998. On publications about scorpions (Arachnida, Scorpiones) by Hemprich and Ehrenberg (1828-1831). Bulletin of the British Arachnological Society 11: 29-35.
- Brignoli, P.M. 1985. On the correct dates of publication of the arachnid taxa described in some works by C.W. Hahn and C.L. Koch (Arachnida). Bulletin of the British Arachnological Society 6: 414-416.
- Brindle, A. 1965. A revision of the subfamily Allostethinae (Dermaptera, Labiduridae). Annals and Magazine of Natural History 13: 575-596.
- Bucherl, W. 1959. Escorpioes e escorpionismo no Brasil. X. Catalogo da colecao escorpionica do Instituto Butantan. Memorias do Instituto de Butantan 29: 255-275.
- Bucherl, W. 1964. Distribucao geografica dos aracnoides peconhentos temiveis. Memorias do Instituto de Butantan 31: 55-66.
- Bucherl, W. 1967. Escorpioes, aranhas e escolopendromorfos da Amazonia. In H. Lent (editor), Atas do simposio sobre a biota amazonica 5 (Zoologia): 111-125.
- Casper, G.S. 1985. Prey capture and stinging behavior in the emperor scorpion, Pandinus imperator (Koch) (Scorpiones, Scorpionidae). Journal of Arachnology 13: 277-283.
- Cekalovic, T.K. 1982. Estado actual de la coleccion aracnologica del Museo de Zoologia de la Universidad de Concepcion (M.Z.U.C.), parte Scorpiones. Memorias do Instituto de Butantan 46: 187-192.
- Chandrasekara Reddy, D., and B. Padmanabhanaidu. 1977. Diurnal rhythmicity of alkaline and acid phosphatases in the scorpion, Heterometrus fulvipes C. Koch. Comparative Physiology and Ecology 2: 217-219.
- Chengal Raju, D., M.D. Bashamohideen, and C. Narasimham. 1973. Circadian rhythm in blood glucose and liver glycogen levels of scorpion, Heterometrus fulvipes. Experientia 29: 964-965.
- Chengal Raju, D., S. Govindappa, and K.S. Swami. 1978. Influence of sex on hepatopancreatic glycogenolysis of scorpion Heterometrus fulvipes (C. Koch). Current Science 47: 111-113.
- Cloudsley-Thompson, J.L. 1981. A comparison of rhythmic locomotory activity in tropical forest Arthropoda with that in desert species. Journal of arid environments 4: 327-334.
- Cloudsley-Thompson, J.L. 1990. Scorpions in mythology, folklore and history. In G.A. Polis (editor), The biology of scorpions: 462-485. Stanford, CA: Stanford University Press.
- Condevaux-Lanloy, C. 1996. Notes sur la parturition d' Heterometrus spinifer (H. & E., 1898) (Scorpiones, Scorpionidae). Arachnides 30: 5-6.
- Copeland, R.J.N. 1990. Notes on the breeding of imperial scorpions (Pandinus imperator). Browse Press.
- Couzijn, H.W.C. 1978. The method of polythetic analysis applied to a source of taxonomic difficulty: the genus Heterometrus H. and E., 1828 (Scorpionidae). Symposium of the Zoological Society of London 42: 327-333.
- Couzijn, H.W.C. 1981. Revision of the genus Heterometrus. Zoologische Verhandelingen 184: 1-196.
- Crucitti, P. 1998 ("1999"). Ricerche bio-ecologiche su Scorpio maurus fuscus (Hemprich and Ehrenberg, 1829) nell'Hatay, Turchia meridionale (Scorpiones, Scorpionidae). Atti della Societa Italiana di Scienzi Naturali (Museo Civico di Storia Naturale di Milano) 139: 203-211.
- Dalingwater, J.E. 1980. SEM observations of the cuticles of some chelicerates. In Proceedings of the 8th Inter- national Congress of Arachnology, Vienna, 1980: 285-289.
- Danin, A. 1994. Association of Salsola inermis and scorpion burrows in leached soils in the Judean desert, Israel. Israel Journal of Plant Sciences 42: 37-40.
- Darriba, D., G.L. Taboada, R. Doallo, and D. Posada. 2012. jModelTest 2: more models, new heuristics and parallel computing. Nature Methods 9: 772.
- Davydov, K.N. 1898. [Notes on some scorpion species of the Palestinian fauna.] Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St.-Petersbourg 3: 39-51. [in Russian]
- DeGeer, C. (K. DeGeer). 1778. Memoires pour servir a l'histoire des insectes. Stockholm: Imprimerie Pierre Hesselberg 7: 1-950.
- DeGeer, C. (K. DeGeer). 1783. Abhandlungen zur Geschichte der Insekten, aus dem Franzosichen uebersetzt und mit Anmerkungen herausgegeben von J.A.E. Goeze. Nurnberg: Gabriel Nikolaus Raspe. Nurnberg 7: 1-275, pl. 4. [translation of DeGeer, 1778]
- Delfosse, E. 1998. Essais de nourriture pour les arachnides. Primere partie: les scorpions (Arachnida, Scorpiones). Arachnides 37: 28-32.
- Devarajulu Naidu, V. 1969. Some studies on the heart beat of scorpion, Heterometrus fulvipes. Experientia 25: 1274.
- Devarajulu Naidu, V., and B. Padmanabhanaidu. 1976. Diurnal rhythmic activity in the heart-beat of the scorpion Heterometrus fulvipes C. Koch. Indian Journal of Experimental Biology 14: 1-5.
- Devarajulu Naidu, V., and S.A.T. Venkatachari. 1974. Succinate dehydrogenase activity in the heart muscles of the scorpion, Heterometrus fulvipes C. Koch. Indian Journal of Experimental Biology 12: 539- 542.
- Di, Z., X. Xu, Z. Cao, Y. Wu, and W. Li. 2013. Notes on the scorpions (Arachnida, Scorpiones) from Xizang with the redescription of Scorpiops jendeki Kovarik, 2000 (Scorpiones, Euscorpiidae) from Yunnan (China). ZooKeys 301: 51-99.
- Dimitrov, D., et al. 2012. Tangled in a sparse spider web: single origin of orb weavers and their spinning work unravelled by denser taxonomic sampling. Proceedings of the Royal Society B, Biological Sciences 279: 1341-1350.
- Dimitrov, D., J.J. Astrin, and B.A. Huber. 2013. Pholcid spider molecular systematics revisited, with new insights into the biogeography and the evolution of the group. Cladistics 29: 132-146.
- Domaniewski, M.J. 1913. Spostrzezenia nad zjawiskami zmiennosci i korrelacyi u niedzwiadka jawanskiego (Heterometrus cyaneus C.L. Koch) (Beitrage zur Kenntnis der Variabilitat und Korrelationserscheinungen bei dem javanischen Skorpion). Bulletin de l'Academie des Sciences de Cracovie (ser. B) 1913: 251-271.
- Dubale, M.S., and A.B. Vyas. 1968. The structure of the chela of Heterometrus sp. and its mode of operation. Bulletin of the Southern California Academy of Sciences 67: 240-244.
- Dufour, L. 1856. Histoire anatomique et physiologique des scorpions. Memoires Presentes par Divers Savants a l'Academie des Sciences de l'Institut Imperial de France 14: 561-653.
- Dumont, H.J. 1982. Relict distribution patterns of aquatic animals: another tool in evaluating Late Pleistocene climate changes in the Sahara and Sahel. Palaeoecology of Africa 14: 1-24.
- Dumortier, B. 1964. Morphology of sound emission apparatus in Arthropoda. In R.-G. Busnel (editor), Acoustic behaviour of animals: 277-345. Amsterdam: Elsevier.
- Dupre, G. 1989. Esquisse de determination du genre Heterometrus H. & E. (Scorpionida: Scorpionidae) d'apres son lieu de capture. Arachnides 3: 4-6.
- Dupre, G. 1990. Le point sur l'elevage de Pandinus (Pandinus) imperator (C.L. Koch 1842) (Scorpionida, Scorpionidae). Arachnides 7: 9-13.
- Dupre, G. 1992. Evolution des importations de scorpions exotiques en France. Arachnides 12: 14-16.
- Dupre, G. 1993. Fiche d'elevage Heterometrus sp. et ssp. Arachnides 16: 13.
- Dupre, G., N. Lambert, and P. Gerard. 1998. Les scorpions: biologie, elevage. Paris: P. Gerard. 82 pp.
- Eastwood, E.B. 1978a. Notes on the scorpion fauna of the Cape. Part 3. Some observations on the distribution and biology of scorpions on Table Mountain. Annals of the South African Museum 74: 229-248.
- Eastwood, E.B. 1978b. Notes on the scorpion fauna of the Cape. Part 4. The burrowing activities of some scorpionids and buthids (Arachnida, Scorpionida). Annals of the South African Museum 74: 249-255.
- Esposito, L.A., H.Y. Yamaguti, C.A. Souza, R. Pinto-da- Rocha, and L. Prendini. 2017. Systematic revision of the Neotropical club-tailed scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, revalidation of Heteroctenus, and descriptions of two new genera and three new species (Buthidae: Rhopalurusinae). Bulletin of the American Museum of Natural History 415: 1-136.
- Esposito, L.A., H.Y. Yamaguti, R. Pinto-da-Rocha, and L. Prendini. 2018. Plucking with the plectrum: phylogeny of the New World buthid scorpion subfamily Centruroidinae Kraus, 1955 (Scorpiones: Buthidae) reveals evolution of three pecten-sternite stridulation organs. Arthropod Systematics and Phylogeny 76: 87-122.
- Fabricius, J.C. 1775. Systema entomologiae. Flensburg et Leipzig: Officina Libraria Kortii. 832 pp.
- Fabricius, J.C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma autorum, loca natalia, metamorphosin adjectis observationibus, descriptionibus 1: 550-552. Bohnii: Impensis Carol. Ernest.
- Fabricius, J.C. 1787. Mantissa insectorum sistens eorum species nuper detectas adjectis characteribus genericis, differentiis, specificis, emendationibus, observationibus, descriptionibus, emendationibus 1. [Scorpiones: p. 348.] Hafniae: Impensis Christ. Gottl. Proft.
- Fabricius, J.C. 1793. Entomologica systematica emendata et aucta, secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus 2: 1-519. Hafniae: Impensis Christ. Gottl. Proft.
- Fage, L. 1933. Les scorpions de l'Indochine francaise, leurs affinites, leurs distribution geographique. Annales de la Societe Entomologique de France 102: 25-34.
- Fage, L. 1936. Nouvelle contribution a l'etude des scorpions de l'Indochine francaise. Bulletin de la Societe Entomologique de France 41: 179-181.
- Fage, L. 1946. Scorpions et pedipalpes de l'Indochine francaise. Annales de la Societe Entomologique de France 113: 71-81.
- Farley, R.D. 2005. Developmental changes in the embryo, pronymph, and first molt of the scorpion Centruroides vittatus (Scorpiones: Buthidae). Journal of Morphology 265: 1-27.
- Farley, R.D. 2008. Development of respiratory structures in embryos and first and second instars of the bark scorpion, Centruroides gracilis (Scorpiones: Buthidae). Journal of Morphology 269: 1134-1156.
- Farley, R.D. 2011. Pectine development in scorpion embryos and first and second instars. Euscorpius 120: 1-47.
- Fet, V. 2000. Family Scorpionidae Latreille, 1802. In V. Fet, W.D. Sissom, G. Lowe, and M.E. Braunwalder, Catalog of the scorpions of the World (1758-1998): 427-486. New York: New York Entomological Society.
- Fet, V., and M.E. Soleglad. 2005. Contributions to scorpion systematics. I. On recent changes in high level taxonomy. Euscorpius 3: 1-13.
- Fitch, W.M. 1971. Toward defining the course of evolution: minimum change for a specific tree topology. Systematic Zoology 20: 406-416.
- Flower, S.S. 1901. Notes on millipedes, centipedes, scorpions, etc. of the Malay Peninsula and Siam. Journal of the Straits Branch of the Royal Asiatic Society 36: 1-48.
- Francke, O.F. 1982. Order Scorpiones. In S.P. Parker (editor), Synopsis and classification of living organisms: 73-75. New York: McGraw Hill.
- Francke, O.F. 1985. Conspectus genericus scorpionorum 1758-1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University 98: 1-32.
- Francke, O.F., and W.D. Sissom. 1984. Comparative review of the methods used to determine the number of molts to maturity in scorpions (Arachnida), with analysis of the post-birth development of Vaejovis coahuilae Williams (Vaejovidae). Journal of Arachnology 12: 1-20.
- Froufe, E., P. Sousa, P.C. Alves, and D.J. Harris. 2008. Genetic diversity within Scorpio maurus (Scorpiones: Scorpionidae) from Morocco: preliminary evidence based on CO1 mitochondrial DNA sequences. Biologia 63: 1157-1160.
- Garnier, G. 1974. Elevage en laboratoire de Pandinus imperator (C.L. Koch, 1842) (Scorpionidae - Scorpioninae). Bulletin de la Societe Zoologique de France 99: 693-699.
- Garnier, G., and R. Stockmann. 1973 ("1971"). Comportement de la reproduction chez le scorpion Pandinus imperator. In Proceedings of the 5th International Congress of Arachnology, Brno, 1971: 13-22.
- Geethabali. 1976. Central course of photic input in the ventral nerve cord of scorpion (Heterometrus fulvipes). Experientia 32: 345-347.
- Geethabali, and K. Pampathi Rao. 1973. A metasomatic neural photoreceptor in the scorpion. Journal of Experimental Biology 58: 189-196.
- Gervais, P.M. 1844a. Scorpions. In C.A. Walckenaer (editor), Histoire naturelle des insectes. Apetres. 3: 14-74. Paris: Librarie Encyclopedique de Roret.
- Gervais, P.M. 1844b. Remarques sur la famille des scorpions et description de plusieurs especes nouvelles de la collection du Museum. Archives du Museum d'Histoire Naturelle 4: 201-240.
- Giltay, M.L. 1931. Scorpions et pedipalpes. Resultats scientifiques du voyage aux Indes Orientales Neerlandaises de LL. AA. RR. le Prince et la Princesse Leopold de Belgique. Memoires du Musee Royal d'Histoire Naturelle de Belgique 3: 1-28.
- Giltay, M.L. 1935. Liste des arachnides d'Extreme-Orient et des Indes Orientales recueillis, en 1932, par S.A.R. le Prince Leopold de Belgique. Bulletin du Musee Royal d'Histoire Naturelle de Belgique 11: 1-3.
- Goloboff, P.A. 1993. Estimating character weights during tree search. Cladistics 9: 83-91.
- Goloboff, P.A., J.S. Farris, and K.C. Nixon. 2003. TNT: tree analysis using new technology. Computer software and documentation. Online resource (https:// cladistics.org/tnt).
- Goloboff, P.A., J.S. Farris, and K.C. Nixon. 2008. TNT, a free program for phylogenetic analysis. Cladistics 24: 774-786.
- Gopalakrishnakone, P., J. Cheah, and M.C.E. Gwee. 1995. Black scorpion (Heterometrus longimanus) as a laboratory animal: maintenance of a colony of scorpion for milking of venom for research, using a restraining device. Laboratory Animals 29: 456-458.
- Gough, L.H., and S. Hirst. 1927. Key to the identification of Egyptian scorpions. A method of identifying Egyptian scorpions from the fifth caudal segment and stings. Bulletin of the Ministry of Agriculture of Egypt, Technical Scientific Service 76: 1-7.
- Govindarajan, S., and G.S. Rajulu. 1974. Presence of resilin in a scorpion Palamnaeus swammerdami and its role in the food-capturing and sound-producing mechanisms. Experientia 30: 908-909.
- Guindon, S., and O. Gascuel. 2003. A simple, fast, and accurate algorithm to estimate large phylogenies by maximum likelihood. Systematic Biology 52: 696-704.
- Harrison, J.L. 1954. A family of the great black scorpion (Heterometrus longimanus). Malayan Nature Journal 9: 31-32.
- Hemprich, F.G., and C.G. Ehrenberg. 1828. Zoologica II. Arachnoidea. Plate I: Buthus; plate II: Androctonus. In Symbolae physicae seu icones et descriptiones animalium evertebratorum sepositis insectis quae ex itinere per Africam borealem et Asiam occidentalem. Decas Prima, Plates IX et X. Berolini: Officina Academica.
- Hemprich, F.G., and C.G. Ehrenberg. 1829. Vorlaufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Scorpione und deren geographischen Verbreitung, nach den eigenen Beobachtungen. Verhandungen der Gesellschaft Naturforschende Freunde in Berlin 1: 348-362.
- Hemprich, F.G., and C.G. Ehrenberg. 1831. Animalia articulata, Arachnoidea. Scorpiones africani et asiatici. In Symbolae physicae. Animalia evertebrata, exclusis insectis. Series prima, tabularum decade prima. Berolini: Officina Academica, 12 pp.
- Henderson, J.R. 1919. Two new scorpions from southern India. Records of the Indian Museum 16: 378-381.
- Herbst, J.F.W. 1800. Naturgeschichte der Skorpionen. In Natursystem der Ungeflugelten Insekten 4: 1-86. Berlin: Gottlieb August Lange.
- Hewitt, J. 1915. New South African Arachnida. Annals of the Natal Museum 3: 289-327.
- Hewitt, J. 1918. A survey of the scorpion fauna of South Africa. Transactions of the Royal Society of South Africa 6: 89-192.
- Hewitt, J. 1925. Facts and theories on the distribution of scorpions in South Africa. Transactions of the Royal Society of South Africa 12: 249-276.
- Hewitt, J. 1931. A new solifuge and scorpion from South-West Africa. Annals of the South African Museum 30: 93-99.
- Hincks, W.D. 1960. Indian Dermaptera of the genus Diplatys Serville (Pygidicranidae), with the description of a new species. Bulletin of Entomology 1: 14-15.
- Hirst, S. 1911. On a collection of Arachnida and Chilopoda made by Mr. S.A. Neave in Rhodesia, north of the Zambezi. Memoirs of the Linnean Philosophical Society 56: 1-11.
- Hjelle, J.T. 1990. Anatomy and morphology. In G.A. Polis (editor), The biology of scorpions: 9-63. Stanford, CA: Stanford University Press.
- Hosoi, F. 1990. [Breeding and display of a scorpion Heterometrus cyaneus.] Insectarium 27: 164-167. [in Japanese]
- Hue, L.-X., Q.-M. Pham, D.-S. Pham, and T.-C. Ngo. 1998. [Scorpions (Scorpionoides) in Vietnam.] Tap Chi Sinh Hoc 20: 7-9. [in Vietnamese]
- Hull-Williams, V. 1986. How to keep scorpions. Fitzgerald Entomological Series 2. London: Fitzgerald Publishing.
- Hull-Williams, V. 1989. A brief description of the collection and appearance of Heterometrus gigantometrus swammerdami titanicus in Sri Lanka. Journal of the British Tarantula Society 5: 12-14.
- Indra, T.J. 2001. Scorpion. In Fauna of Nilgiri Biosphere Reserve. Zoological Survey of India, Fauna of Conservation Area Series II: 55-58.
- Indra, T.J. 2006. Scorpion. In Fauna of Bilgiri Rangaswamy Temple Wildlife Sanctuary. Zoological Survey of India, Conservation Area Series 27: 197-200.
- Indra, T.J. 2009. Scorpionida. In Fauna of Tamil Nadu. Zoological Survey of India, Fauna Series 17: 141-143.
- IUCN (International Union for the Conservation of Nature). 1994. Analyses of proposals to amend the CITES appendices. Inclusion of Emperor Scorpions Pandinus dictator, P. gambiensis and P. imperator in appendix II. Ghana. IUCN Document 9.47.63-65: 186-188.
- Janion-Scheepers, C., et al. 2016. Soil biota in a megadiverse country: current knowledge and future directions in South Africa. Pedobiologia - Journal of Soil Ecology 59: 129-174.
- Javed, S.M.M., Z.A. Mirza, F. Tampal, and W.R. Lourenco. 2010a. A new species of the genus Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from India with notes on its natural history. Boletin de la Sociedad Entomologica Aragonesa 47: 143- 148.
- Javed, S.M.M., Z.A. Mirza, R.V. Sanap, and F. Tampal. 2010b. First record of Liocheles nigripes Pocock, 1897 (Scorpiones: Hemiscorpiidae) from Andhra Pradesh with a checklist of scorpions of the state. Journal of Threatened Taxa 2: 783-785.
- Jayaram, V., and B. Padmanabhanaidu. 1979. Carbohydrate metabolism in the heart of the scorpion, Heterometrus fulvipes (C.L. Koch) with reference to sex. Current Science 48: 6-8.
- Jayaram, V., and B. Padmanabhanaidu. 1980. Utilisation of carbohydrates in the heart of the scorpion Heterometrus fulvipes (C.L. Koch). Proceedings: Animal Sciences 89: 109-112.
- Jayaram, V., D. Chandrasekara Reddy, and B. Padmanabhanaidu. 1978. Circadian rhythmicity in phosphorylase activity and glycogen content in the heart muscle of the scorpion, Heterometrus fulvipes, C. L. Koch. Experientia 34: 1184-1185.
- Kalarani, V., P. Murali Mohan, and R.W. Davies. 1992. Thermal acclimation and metabolism of the hepatopancreas in the tropical scorpion, Heterometrus fulvipes. Journal of Thermal Biology 17: 141-146
- Kamenz, C., and L. Prendini. 2008. An atlas of book lung ultrastructure in the order Scorpiones (Arachnida). Bulletin of the American Museum of Natural History 316: 1-259.
- Kanter, H. 1971. Giftschlangen und Skorpione Nordafrikas. In H. Schiffers (editor), Die Sahara und ihre Randgebiete: 587-603. Munich: Weltforum Verlag.
- Kanungo, M.S. 1954. The ecology and behaviour of the scorpion Palamnaeus bengalensis C. Koch. Journal of the Bombay Natural History Society 52: 536-539.
- Karsch, F. 1879a. Skorpionologische Beitrage. Part I. Mitteilungen des Munchener Entomologischen Vereins 3: 6-22.
- Karsch, F. 1879b. Skorpionologische Beitrage. Part II. Mitteilungen des Munchener Entomologischen Vereins 3: 97-136.
- Karsch, F. 1884. Ueber einige neue und minder bekannte Arthropoden des Bremer Museums. Abhandlungen, Herausgegeben vom Naturwissenschaftlichen Verein zu Bremen 9: 65-71.
- Karsch, F. 1892. Arachniden von Ceylon und von Minikoy, gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift 36: 267-310.
- Kastner, A. 1931. Die Hufte und ihre Umformung zu Mundwerkzeugen bei den Arachniden. Zeitschrift fur Morphologie und Okologie der Tiere 22: 721-758.
- Kastner, A. 1941. 1. Ordnung der Arachnida: Scorpiones. In T. Krumbach (editor), Handbuch der Zoologie 3: 117-240. Berlin: Walter de Gruyter Verlag.
- Katoh, K., and D.M. Standley. 2013. MAFFT multiple sequence alignment software version 7: improvements in performance and usability. Molecular Biology and Evolution 30: 772-780.
- Katoh, K., K. Misawa, K.I. Kuma, and T. Miyata. 2002. MAFFT: a novel method for rapid multiple sequence alignment based on fast Fourier transform. Nucleic Acids Research 30: 3059-3066.
- Keyserling, E. 1885. Die Arachniden Australiens. Ordo Scorpiones. In Die Arachniden Australiens nach der Natur beschrieben und abgebildet begonnen von Dr. L. Koch 2: 1-86, tab. I-VI. Nurnberg: Bauer and Raspe.
- Khatavkar, R.S., and N.K. More. 1990. A note on the burrow and burrowing activity of Heterometrus xanthopus Pocock (Arachnida: Scorpionidae). Journal Soil Biology and Ecology 10: 79-81.
- Khatoon, S. 1999. Scorpions of Pakistan (Arachnida: Scorpionida). Proceedings of the Pakistan Congress of Zoology 19: 207-225.
- Kishida, K. 1939. Arachnida of Jehol. Order Scorpiones. Report of the first scientific expedition to Manchoukuo under the leadership of Shigeyasu Tokunaga, June-October, 1933, Sect. 5, 1(4) 10: 1-67.
- Klussmann-Fricke, B.-J., L. Prendini, and C.S. Wirkner. 2012. Evolutionary morphology of the hemolymph vascular system in scorpions: a character analysis. Arthropod Structure and Development 41: 545-560.
- Koch, C.L. 1836. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 3 (1-5): 1-104, pl. 73-105 (figs. 164-245).
- Koch, C.L. 1837. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 4 (1-5): 1-108, pl. 119-138 (figs. 253-318).
- Koch, C.L. 1838. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 4 (6): 109-144, pl. 139-144 (figs. 319-339).
- Koch, C.L. 1840. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 8 (1, 2): 1-40, pl. 248-264 (figs. 595-620).
- Koch, C.L. 1841a. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 8 (3-6): 41-131, pl. 265- 288 (figs. 621-694).
- Koch, C.L. 1841b. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 9 (1): 1-56, pl. 289-306 (figs. 695-726).
- Koch, C.L. 1850. Scorpionen. In Ubersicht des Arachnidensystems. Nurnberg: C.H. Zeh'sche Buchhandlung 5: 86-92.
- Koch, L.E. 1977. The taxonomy, geographic distribution and evolutionary radiation of Australo-Papuan scorpions. Records of the Western Australian Museum 5: 83-367.
- Kopstein, F. 1921. Die Skorpione des Indo-Australischen Archipels. Mit Grundlage der in hollandischen Sammlungen, vornahmlich des Rijks-Museums in Leiden, vorhandenen Arten. Zoologische Mededeelingen 6: 115-144.
- Kopstein, F. 1923. Liste der Skorpione des Indo-Australischen Archipels im Museum zu Buitenzorg. Treubia 3: 184-187.
- Kopstein, F. 1926. De Schorpioenen van Java. De Troopischer Natuur 15: 109-118.
- Kopstein, F. 1927. Der javanische Reisenskorpion. Der Zoologische Reisdoor Tropen 1927: 102-108.
- Kopstein, F. 1929. Spolia Mentawiensia. Scorpiones. Bulletin of the Raffles Museum 2: 124.
- Kotzman, M., Y. Lubin, and M. Goldberg. 1989. The activities of a burrow-dwelling scorpion revealed at the surface. Acta Zoologica Fennica 190: 205- 208.
- Kovarik, F. 1992. A check list of scorpions (Arachnida: Scorpiones) in the collections of the Zoological Department, National Museum in Prague. Acta Societatis Zoologicae Bohemicae 56: 181-186.
- Kovarik, F. 1994. Isometrus zideki sp. n. from Malaysia and Indonesia, and a taxonomic position of Isometrus formosus, I. thurstoni and I. sankariensis (Arachnida: Scorpionida: Buthidae). Acta Societatis Zoologicae Bohemicae 58: 195-203.
- Kovarik, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae 59: 187-207.
- Kovarik, F. 1997. A check-list of scorpions (Arachnida) in the collections of the Hungarian Natural History Museum, Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 89: 177-185.
- Kovarik, F. 1998. Stiri [Scorpions]. Jihlava, Czech Republic: Madagaskar, 176 pp. [in Czech]
- Kovarik, F. 2000. Pandinus (Pandinops) pococki sp. n. from Somalia, and Pandinus pugilator, a junior synonym of Pandinus (Pandinops) bellicosus comb. n. (Scorpiones, Scorpionidae). Serket 7: 1-7.
- Kovarik, F. 2002. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany. Serket 8: 1-23.
- Kovarik, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones), with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae 67: 133-159.
- Kovarik, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius 15: 1-60.
- Kovarik, F. 2009. Illustrated catalog of scorpions. Part 1. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clarion Production. 170 pp.
- Kovarik, F. 2011. A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae). Euscorpius 129: 1-18.
- Kovarik, F. 2012. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius 141: 1-22.
- Kovarik, F. 2013. Pandinus (Pandinus) trailini sp. n. from Ethiopia (Scorpiones: Scorpionidae), with data on localities and life strategy. Euscorpius 163: 1-14.
- Kovarik, F. 2016. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VIII. Pandinops Birula, 1913 (Scorpionidae), with description of two new species. Euscorpius 229: 1-20.
- Kovarik, F. 2017. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XI. Pandinurus kmoniceki sp. n. (Scorpionidae) from Somaliland. Euscorpius 243: 1-14.
- Kovarik, F., et al. 2016. Scorpions of Sri Lanka (Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat. n. Euscorpius 220: 1-133.
- Kovarik, F., S. Navidpour, and M.E. Soleglad. 2017a. Hemiscorpius shahii sp. n. from Iran (Scorpiones: Hemiscorpiidae). Euscorpius 249: 1-9.
- Kovarik, F., G. Lowe, M.E. Soleglad, and J. Pliskova. 2017b. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part X. Pandiborellius stat. n. and Pandinurus (Scorpionidae) with description of four new species from Eritrea and Ethiopia, and review of Pandinus sensu lato taxonomy. Euscorpius 238: 1-103.
- Kovarik, F., et al. 2017c. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XII. Pandinurus hangarale sp. n. (Scorpionidae) from Somaliland and a review of type locality and true distribution of Pandinurus smithi (Pocock, 1897). Euscorpius 253: 1-18.
- Kovarik, F., et al. 2019a. Scorpions of Sri Lanka (Arachnida: Scorpiones). Part III. Heterometrus yaleensis sp. n. (Scorpionidae). Euscorpius 283: 1-13.
- Kovarik, F., G. Lowe, and T. Mazuch. 2019b. Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XIX. Pandiborellius meidensis (Karsch, 1879) and Pandinurus fulvipes sp. n. (Scorpionidae) from Somaliland. Euscorpius 275: 1-18.
- Kraepelin, K. 1894. Revision der Skorpione. II. Scorpionidae und Bothriuridae. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 11: 1-248.
- Kraepelin, K. 1897. Scorpione und Thelyphoniden. In Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo, im Auftrage der Senckenbergischen naturforschenden Gesellschaft ausgefuhrt von Dr. Willy Kukenthal. Abhandlungen de Senckenbergischen Naturforschenden Gesellschaft 23: 537.
- Kraepelin, K. 1898. Uber die Linneschen Arten der Gattung Scorpio. Zoologischer Anzeiger 564-565: 438-447.
- Kraepelin, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (editor), Das Tierreich, 8 (Arachnoidea): 1-265. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag.
- Kraepelin, K. 1901. Catalogue des scorpions des collections du Museum d'Histoire Naturelle de Paris. Bulletin du Museum National d'Histoire Naturelle, Paris 7: 265-274.
- Kraepelin, K. 1904. Zur Nomenklatur der Skorpione und Pedipalpen. Zoologischer Anzeiger 28: 195-204.
- Kraepelin, K. 1905. Die geographische Verbreitung der Scorpione. Zoologische Jahrbucher, Abtheilung fur Systematik 22: 321-364.
- Kraepelin, K. 1908 ("1907"). Die sekundaren Geschlechtscharaktere der Skorpione, Pedipalpen und Solifugen. Mitteilungen aus dem Naturhistorischen Museum. (2. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten) 25: 181-225.
- Kraepelin, K. 1913 ("1912"). Neue Beitrage zur Systematik der Gliederspinnen. III. A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch-Ostafrikas. Mitteilungen aus dem Naturhistorischen Museum. (2. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten) 30: 123-196.
- Krapf, D. 1988. Skorpione II. Fortpflanzung, Wachstum und Haltung. Herpetofauna (Weinstadt) 56: 24-33.
- Kriesch, M. 1994. Note sur la sociabilite du scorpion Pandinus imperator (C.L. Koch, 1842). Arachnides 21: 9.
- Lamoral, B.H. 1972. New and little known scorpions, solifuges and spiders from the Namib Desert, S.W.A. Madoqua (II) 1: 117-131.
- Lamoral, B.H. 1978a. Systematics and bionomics of the scorpions of South West Africa (Arachnida: Scorpionida). Ph.D. dissertation, University of Natal, Pietermaritzburg, South Africa.
- Lamoral, B.H. 1978b. Soil hardness, an important and limiting factor in burrowing scorpions of the genus Opisthophthalmus C.L. Koch, 1837 (Scorpionidae, Scorpionida). Symposium of the Zoological Society of London 42: 171-181.
- Lamoral, B.H. 1979. The scorpions of Namibia (Arachnida: Scorpionida). Annals of the Natal Museum 23: 497-784.
- Lamoral, B.H. 1980. A reappraisal of the suprageneric classification of Recent scorpions and their zoogeography. In J. Grube (editor), Verhandlungen. 8. Internationaler Arachnologen-Kongress abgehalten an der Universitat fur Bodenkultur Wien, 7-12 Juli, 1980: 439-444. Vienna: H. Egermann.
- Lamoral, B.H., and S.C. Reynders. 1975. A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973. Annals of the Natal Museum 22: 489-576.
- Lampe, E. 1918. Katalog der Skorpione, Pedipalpen und Solifugen des Naturhistorischen Museums der Residenzstadt Wiesbaden. Jahrbucher des Nassauischen Vereins fur Naturkunde 70: 185-203.
- Lankester, E.R. 1885. Notes on certain points of the anatomy of generic characters of scorpions. Transactions of the Zoological Society, London 11: 372-384.
- Larrouy, G., M.C. Signorel, and Y. Cambefort. 1973. Comportement en captivite de Pandinus imperator C.L. Koch et connaissance des jeunes. Bulletin de la Societe d'Histoire Naturelle de Toulouse 109: 346- 350.
- Latreille, P.A. 1802 ("An X"). Histoire naturelle, generale et particuliere, des crustaces et des insectes. 3. (Ouvrage faisant suite a l'histoire naturelle generale et particuliere, composee par Leclerc de Buffon, et redigee par C.S. Sonnini.) Paris: F. Dufart.
- Latreille, P.A. 1803. Scorpion. In Nouveau dictionnaire d'histoire naturelle, appliquee aux arts 20: 264-268.
- Latreille, P.A. 1804. Histoire des scorpions. In Histoire naturelle, generale et particuliere, des crustaces et des insectes. Ouvrage faisant suite aux Oeuvres de Leclerc de Buffon, et partie de cours complet d'histoire naturelle redigee par C.S. Sonnini, 7: 111- 129. Paris: F. Dufart.
- Latreille, P.A. 1806. Genera crustaceorum et insectorum secundum ordinem naturalem in familias dis- posita, iconibus exemplisque plurimis explicata. Tome 1. Paris: Amand Koenig, 302 pp. [Scorpiones: pp. 130-132.]
- Latreille, P.A. 1810. Considerations generales sur l'ordre naturel des animaux composant les classes des crustaces, des arachnides et des insectes. Paris: F. Schoell, 444 pp. [Scorpiones: pp. 116, 118.]
- Latreille, P.A. 1817. Les crustaces, les arachnides et les insectes. In [G.G.L.C.F.D.] Cuvier (editor), Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee. 3. Paris: Deterville.
- Latreille, P.A. 1825. Familles naturelles du regne animal, exposees succinctement et dans un ordre analytique, avec l'indication de leurs genres. Paris: J.B. Bailliere.
- Laurie, M. 1890. The embryology of the scorpion (Euscorpius italicus). Quarterly Journal of Microscopical Science, New Ser. 31: 105-141, pl. XIII-XVIII.
- Laurie, M. 1891. Some points in the development of Scorpio fulvipes. Quarterly Journal of Microscopical Science, New Ser. 32: 587-597, pl. XL.
- Laurie, M. 1896a. Notes on the anatomy of some scorpions, and its bearing on the classification of the order. Annals and Magazine of Natural History (Ser. 6) 17: 185-193.
- Laurie, M. 1896b. Further notes on the anatomy and development of scorpions, and their bearing on the classification of the order. Annals and Magazine of Natural History (Ser. 6) 17: 121-133.
- Laverack, M.S. 1966. Observations on a proprioceptive system in the legs of the scorpion Hadrurus hirsutus. Comparative Biochemistry and Physiology 19: 241- 251.
- Lawrence, R.F. 1928. Contributions to a knowledge of the fauna of South-West Africa. VII. Arachnida (Part 2). Annals of the South African Museum 25: 217-312.
- Lawrence, R.F. 1969. A new genus of psammophile scorpion and new species of Opisthophthalmus from the Namib Desert. Scientific Papers of the Namib Desert Research Station 48: 105-116.
- Lawrence, R.F. 1971. Spiders and other Arachnida. In D.J. Potgieter, P.C. du Plessis, and S.H. Skaife. Animal life in southern Africa: 210-217. Cape Town: NASOU Ltd.
- Lawrence, R.F. 1973. Scorpions. In J.J. Spies and P.C. du Plessis (editors), Standard encyclopaedia of southern Africa: 543-544. Cape Town: NASOU Ltd. [Leach, W.E.] 1814. Scorpionides. In D. Brewster (editor), Edinburgh encyclopaedia 7: 412. Edinburgh: William Blackwoods and Sons. [anonymous article]
- Levy, G., and P. Amitai. 1980. Fauna Palaestina. Arachnida. I. Scorpiones. Jerusalem: Israel Academy of Sciences and Humanities.
- Lewis, P.O. 2001. A likelihood approach to estimating phylogeny from discrete morphological character data. Systematic Biology 50: 913-925.
- Linnaeus, C. (C. von Linne). 1746. Museum Adolpho- Fridericianum, dissertatio, respondens Laur. Balk. 4-48 pp. Holmiae (Stockholm).
- Linnaeus, C. (C. von Linne). 1748. Systema naturae sistens regna tria naturae, in classes et ordines, genera et species redacta tabulisque aeneis illustrata. Ed. 6, emendata et aucta. 1: 1-224. [Scorpiones: p. 68.] Holmiae (Stockholm): Gododfr. Kiesewetteri.
- Linnaeus, C. (C. von Linne). 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, loci. Ed. 10. 1: 1-821. [Scorpiones: pp. 624, 625.] Holmiae (Stockholm): Laurentii Salvii.
- Linnaeus, C. (C. von Linne). 1767. Systema naturae per regna tria naturae, secudum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Ed. 12. 1: 533-1327. Holmiae (Stockholm): Laurentii Salvii.
- Linnaeus, C. (C. von Linne). 1775. Scorpionen. In Natursystem, tab. 33: 1088-1092.
- Lonnberg, E. 1897a. Om skorpionernas och pedipalpernas geografiska utbredning. Entomologisk Tidskrift 18: 193-211.
- Lonnberg, E. 1897b. Scorpioner och pedipalper i Upsala Universitets Zoologiska Museum. Entomologisk Tidskrift 18: 175-192.
- Lonnberg, E. 1898a. A revision of the Linnean type specimens of scorpions and pedipalps in the Zoological Museum of the Royal University of Upsala. Annals and Magazine of Natural History 7: 82-89.
- Lonnberg, E. 1898b. Noch einmal uber die Linne'schen Arten der Gattung Scorpio. Zoologischer Anzeiger 21: 549-552.
- Loria, S.F., and L. Prendini. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS One 9: e112913. [doi: 10.1371/journal.pone.0112913]
- Loria, S.F., and Prendini, L. (in press). Burrowing into the forest: phylogeny of the Asian forest scorpions (Scorpionidae: Heterometrinae) and the evolution of ecomorphotypes. Cladistics.
- Lourenco, W.R. 1989. Retablissement de la famille des Ischnuridae, distincte des Scorpionidae Pocock, 1893, a partir de la sous-famille des Ischnurinae, Pocock, 1893. Revue Arachnologique 8: 159-177.
- Lourenco, W.R. 1999. Considerations taxonomiques sur le genre Hadogenes Kraepelin, 1894; creation de la sous-famille des Hadogeninae n. subfam., et description d'une espece nouvelle pour l'Angola (Scorpiones, Scorpionidae, Hadogeninae). Revue Suisse de Zoologie 106: 929-938.
- Lourenco, W.R. 2000. Panbiogeographie, les familles des scorpions et leur repartition geographique. Biogeographica 76: 21-39.
- Lourenco, W.R. 2002. Reproduction in scorpions, with special reference to parthenogenesis. In S. Toft and N. Scharff (editors), European Arachnology 2000: 71-85. Aarhus: Aarhus University Press.
- Lourenco, W.R. 2009. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-Saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 15: 99-113.
- Lourenco, W.R. 2014. Further considerations on the identity and distribution of Pandinus imperator (C.L. Koch, 1841) and description of a new species from Cameroon (Scorpiones: Scorpionidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 17: 139-151.
- Lourenco, W.R., and J.L. Cloudsley-Thompson. 1996. Recognition and distribution of the scorpions of the genus Pandinus Thorell, 1876 accorded protection by the Washington Convention. Biogeographica 72: 133-143.
- Lourenco, W.R., and J.L. Cloudsley Thompson. 2009. A new species of the genus Scorpio Linnaeus 1758 from Sudan (Scorpiones, Scorpionidae). Boletin de la Sociedad Entomologica Aragonesa 45: 123-126.
- Lourenco, W.R., and J.L. Cloudsley Thompson. 2012. About the enigmatic presence of the genus Scorpio Linnaeus, 1758 in Congo with the description of a new species from Niger (Scorpiones, Scorpionidae). Serket 13: 1-7.
- Lourenco, W.R., and D. Huber. 2002. New addition to the scorpion fauna (Arachnida: Scorpiones) of Sri Lanka. Revue Suisse de Zoologie 109: 265-275.
- Lourenco, W.R., and A. Rossi. 2016. Confirmation of a new species of Scorpio Linnaeus, 1758 in the Tassili n'Ajjer Mountains, south Algeria (Scorpiones: Scorpionidae). Onychium 12: 11-18.
- Lourenco, W.R., J.-X. Qi, and M.-S. Zhu. 2005. Description of two new species of scorpions from China (Tibet) belonging to the genera Mesobuthus Vachon (Buthidae) and Heterometrus Ehrenberg (Scorpionidae) Zootaxa 985: 1-16.
- Lourenco, W.R., B. Duhem, and J.L. Cloudsley Thompson. 2012. Scorpions from Ennedi, Kapka and Tibesti, the mountains of Chad, with descriptions of nine new species (Scorpiones: Buthidae, Scorpionidae). Arthropoda Selecta 21: 307-338.
- Mahsberg, D. 1990. Brood care and family cohesion in the tropical scorpion Pandinus imperator (Koch) (Scorpiones: Scorpionidae). Acta Zoologica Fennica 190: 267-272.
- Mahsberg, D., R. Lippe, and S. Kallas. 1999. Skorpione. Munster: Natur und Tier Verlag.
- Majumder, S.C., and A. Dey. 2005. Studies on some enthnomedicinal arachnids and insects in relation to their usage as drugs among the tribals of Sundarbans, West Bengal, India. Records of Zoological Survey of India, Occasional Paper 236: 1-38.
- Makioka, T. 1992a. Reproductive biology of the viviparous scorpion, Liocheles australasiae (Fabricius) (Arachnida, Scorpiones, Ischnuridae) III. Structural types and functional phases of the adult ovary. Invertebrate Reproduction and Development 21: 207-214.
- Makioka, T. 1992b. Reproductive biology of the viviparous scorpion, Liocheles australasiae (Fabricius) (Arachnida, Scorpiones, Ischnuridae) II. Repeated pregnancies in virgins. Invertebrate Reproduction and Development 21: 161-167.
- Marx, G. 1890 ("1889"). Arachnida. In L.O. Howard (editor), Scientific results of the explorations by the U.S. Fish Commission Steamer Albatross. No. V. - Annotated catalogue of the insects collected in 1887-'88. Proceedings of the United States National Museum 12: 207-211.
- Masi, L. 1912. Note sugli scorpioni appartenenti al R. Museo Zoologico di Roma. Bollettino Societa Zoologica Italiana 1: 88-108, 120-144.
- Mastanaiah, S., D. Chengul Raju, and K.S. Swami. 1977. Rhythmic variation in the isocitrate dehydrogenase activity in the scorpion Heterometrus fulvipes (C. Koch). Experientia 33: 1051-1052.
- Mastanaiah, S., V. Doraswamy Reddy, D. Chengal Raju, and K.S. Swami. 1978. Circadian rhythmic activity of lipase in the scorpion, Heterometrus fulvipes (C. Koch). Current Science 47: 130-132.
- Mastanaiah, S., D. Chengul Raju, D. Ravi Varma, and K.S. Swami. 1979. Rhythmic variations in the activities of aldolase and isocitrate dehydrogenase in the heart muscle of the scorpion, Heterometrus fulvipes (C. Koch). Experientia 35: 69-70.
- Mathew, A.P. 1948. Nutrition in the advanced scorpion embryos: Palamnaeus scaber Thorell. Proceedings of Indian Academy of Sciences, Section B 27: 111-118.
- Mathew, A.P. 1956. Embryology of Heterometrus scaber. Bulletin of the Central Research Institute (University of Travancore, India) 1: 1-96.
- Mathew, A.P. 1957. Mating in scorpions. Journal of the Bombay Natural History Society 54: 853-857.
- Mathew, A.P. 1962. Changes in the structure of the ovarian and diverticular mucosa in Heterometrus scaber (Thorell) during a reproductive cycle. In R.V. Seshaiya and B.S. Chauhan (editors), Proceedings of the First All-India Congress of Zoology, 24-28 October 1959, University of Jabalpur, Part 2, pp. 100-111. Calcutta: Zoological Society of India.
- Mathew, A.P. 1965. On the movable claw of the pedipalp in the scorpion Heterometrus scaber. Journal of Animal Morphology and Physiology 12: 271-275.
- Mathew, A.P. 1966. The movable claw of the pedipalp of the scorpion, Heterometrus scaber. In G.S. Thapar (editor), Proceedings of the Second All-India Congress of Zoology, October 1962, Banaras Hindu University, Varanasi, Part 2, pp. 411-414. Calcutta: Zoological Society of India.
- Mathew, A.P. 1968. Embryonic nutrition in Urodacus abruptus Pocock. Journal of Animal Morphology and Physiology 15: 162-167.
- McCormick, S.J., and G.A. Polis. 1990. Prey, predators and parasites. In G.A. Polis (editor), The biology of scorpions: 294-320. Stanford, CA: Stanford University Press.
- Meise, W. 1932. Fehlender und extrem entwickelter Sexualdimorphismus im Formenkreis Heterometrus longimanus (Scorpion). Archiv fur Naturgeschichte, Neue Folge 1: 660-671.
- Menzel, P., and F. D'Aluisio. 1998. Man eating bugs: the art and science of eating insects. Berkeley, CA: Material World Books/Ten Speed Press.
- Michalik, P., and D. Mercati. 2010. First investigation of the spermatozoa of a species of the superfamily Scorpionoidea (Opistophthalmus penrithorum, Scorpionidae) with a revision of the evolutionary and phylogenetic implications of sperm structures in scorpions (Chelicerata, Scorpiones). Journal of Zoological Systematics and Evolutionary Research 48: 89-101.
- Miles, C. 2015. The earwig collection (Dermaptera) of the Manchester Museum, U.K., with a complete type catalogue. European Journal of Taxonomy 141: 1-138.
- Millot, J., and M. Vachon. 1949. Ordre des scorpions. In P.-P. Grasse (editor), Traite de zoologie, vol. 6: 386-436. Paris: Masson et Cie.
- Mirza, Z.A., and R. Sanap. 2009. Notes on the reproductive biology of Heterometrus phipsoni Pocock, 1893 (Scorpiones, Scorpionidae). Journal of Threatened Taxa 1: 488-490.
- Mirza, Z.A., D. Joshi, G. Desouza, and R.V. Sanap. 2012. Description of a new species of scorpion of the genus Heterometrus Ehrenberg, 1828 (Scorpiones, Scorpionidae) from the Western Ghats, India. Indian Journal of Arachnology 1: 1-8.
- Monod, L., and W.R. Lourenco. 2005. Hemiscorpiidae (Scorpiones) from Iran, with descriptions of two new species and notes on biogeography and phylogenetic relationships. Revue Suisse de Zoologie 112: 869-942.
- Monod, L., and L. Prendini. 2014. Evidence for Eurogondwana: the roles of dispersal, extinction and vicariance in the evolution and biogeography of Indo-Pacific Hormuridae (Scorpiones: Scorpionoidea). Cladistics 31: 71-111.
- Montambaux, P. 1996. Maintenance en captivite de Pandinus imperator. Reptile Magazine 5: 15-16.
- Moritz, M., and S.C. Fischer. 1980. Die Typen der Arachniden-Sammlung des Zoologischen Museums Berlin. III. Scorpiones. Mitteilungen aus dem Zoologischen Museum in Berlin 56: 309-326.
- Nagaraj, S.K., P. Dattatreya, and T.N. Boramuthi, 2015. Indian scorpions collected in Karnataka: maintenance in captivity, venom extraction and toxicity studies. Journal of Venomous Animals and Toxins including Tropical Diseases 21: 51. [doi: 10.1186/ s40409-015-0053-4]
- Nair, B., and P.A. Kurup. 1975. Investigations on the venom of the Indian scorpion Heterometrus scaber. Biochimica et Biophysica Acta 381: 165-174.
- Nayar, K.K. 1966. Endocrine nature of the lymphoid organs of the scorpion, Heterometrus scaber Thor. In G.S. Thapar (editor), Proceedings of the Second All- India Congress of Zoology, October 1962, Banaras Hindu University, Varanasi, Part 2, pp. 173-174.
- Nemenz, H., and J. Gruber. 1967. Experimente und Beobachtungen an Heterometrus longimanus petersii (Thorell) (Scorpiones). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 107: 5-24.
- Nenilin, A.B., and V. Fet. 1992. Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones). Arthropoda Selecta 1: 3-31. [in Russian; extended English summary]
- Newlands, G. 1972a. Notes on psammophilous scorpions and a description of a new species (Arachnida: Scorpionida). Annals of the Transvaal Museum 27: 241-257.
- Newlands, G. 1972b. Ecological adaptations of Kruger National Park scorpionids (Arachnida: Scorpiones). Koedoe 15: 37-48.
- Newlands, G. 1978. Arachnida (except Acari). In M.J.A. Werger (editor), Biogeography and ecology of southern Africa 2: 677-684. The Hague: W. Junk.
- Newlands, G. 1987. Scorpions. Pretoria: De Jager-HAUM.
- Newport, G. 1843. On the structure, relations, and development of the nervous and circulatory systems, and on the existence of a complete circulation of the blood in vessels, in Myriapoda and macrourous Arachnida. Philosopical Transactions of the Royal Society of London 133: 243-302.
- Niethammer, G. 1971. Die Fauna der Sahara. In H. Schiffers (editor), Die Sahara und ihre Randgebiete: 499-587. Munich: Weltforum Verlag.
- Nixon, K.C. 2002. WinClada, ver. 1.00.08. Computer software and documentation. Online resource (http://www.diversityoflife.org/winclada).
- Pallary, P. 1929. Les scorpions du Sahara central. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord 20: 133-141.
- Pallary, P. 1934. Scorpions du Sahara central. Mission du Hoggar III (fevrier a mai 1928). Memoires de la Societe d'Histoire Naturelle de l'Afrique du Nord 4: 90-100.
- Pallary, P. 1938. Sur les scorpions de la Berberie, de la Syrie et du Congo. Archives de l'Institut Pasteur d'Algerie 16: 279-282.
- Pampapathi Rao, K. 1964. Neurophysiological studies of an arachnid, the scorpion Heterometrus fulvipes. Journal of Animal Morphology and Physiology 11: 133-142.
- Pampapathi Rao, K., and M. Habibulla. 1973. Correlation between neurosecretion and some physiological functions of the scorpion Heterometrus swammerdami. Proceedings of the Indian Academy of Sciences, Section B 77: 148-155.
- Pampapathi Rao, K., and V.S.R. Murthy. 1966. Oscillographic analysis of some proprioceptors in the scorpion Heterometrus fulvipes. Nature 212: 520-521.
- Pande, S., D. Bastawade, A. Padhye, and A. Pawashe. 2012. Diversity of scorpion fauna of Saswad-Jejuri, Pune District, Maharashtra, western India. Journal of Threatened Taxa 4: 2381-2389.
- Patil, A.E., and U.H. Shah. 2012a. Hemocytes in scorpion - Heterometrus xanthopus. Bioscan 7: 139-141.
- Patil, A.E., and U.H. Shah. 2012b. Ultrastructure studies of hemocytes in scorpion Heterometrus xanthopus. Bioscan 7: 491-493.
- Patil, A.E., and U.H. Shah. 2013. Hemocytes in scorpion - Heterometrus phipsoni. Bioscan 8: 33-35.
- Pavesi, P. 1880. Studi sugli aracnidi africani. I. Aracnidi di Tunisia. Annali del Museo Civico di Storia Naturale di Genova 15: 283-388.
- Pavesi, P. 1884. Materiali per lo studio della fauna Tunisina raccolta da G. e L. Doria: aracnidi. Annali del Museo Civico di Storia Naturale di Genova 20: 446-486.
- Pavesi, P. 1895a. Viaggio del Dott. E. Festa in Palestina, nel Libano e regioni vicine. XIV. Aracnidi. Bollettino dei Musei di Zoologia ed Anatomia Comparata 10: 1-11.
- Pavesi, P. 1895b. Aracnidi raccolti ne paese dei Somali dall'ing L. Bricchetti-Robecchi. Bollettino Scientifico, Pavia 17: 37-46.
- Pavesi, P. 1898. Sugli aracnidi raccolti a Giava dal dott. Penzig nel 1895-96. Bollettino Scientifico, Pavia 20: 93-96.
- Pavlovsky, E.N. 1924a. On the morphology of the male genital apparatus in scorpions. Trudy Leningradskogo Obshchestva Yestestvoispytatelei [Transactions of the Leningrad Society of Naturalists] 53: 17-86.
- Pavlovsky, E.N. 1924b. Studies on the organization and development of scorpions. (Introduction. 1: On the external morphology of the scorpions. 2: The integument and its processes. 3: The circulatory system. 4: The system of lymphatic/phagocytic organs). Quarterly Journal of Microscopical Science 68: 615-640.
- Pavlovsky, E.N. 1925. Zur Morphologie des weiblichen Genitalapparatus und zur Embryologie der Skorpione. Annuaire du Musee Zoologique de l'Academie des Sciences d'URSS 26: 137-205.
- Perez Minnocci, S. 1974. Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identificacion de los generos de la region Paleartica Occidental. Madrid: Universidad Complutense de Madrid, Facultad de Ciencias, Departamento de Zoologia, Catedra de Artropodos 7: 1-45.
- Peters, W. 1861. [Ueber eine neue Eintheilung der Skorpione und ueber die von ihm in Mossambique gesammelten Arten von Skorpionen.] Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin 1861: 507-516.
- Petrunkevitch, A. 1955. Arachnida. In R.C. Moore (editor), Treatise on invertebrate paleontology. Part P. Arthropoda. 2. Chelicerata with sections on Pycnogonida and Palaeoisopus: 42-162. Lawrence, KS: Geological Society of America and University of Kansas Press.
- Pham, D.S., T.H. Tran, and W.R. Lourenco. 2017. Diversity and endemicity in the scorpion fauna of Vietnam. A preliminary synopsis. Comptes Rendus Biologies 340: 132-137.
- Pickford, M., and B. Senut. 2000. Geology and palaeobiology of the central and southern Namib Desert, southwestern Africa. Vol. 1: Geology and history of study. Geological Survey of Namibia, Memoir 18: 1-155. Windhoek: Ministry of Mines and Energy.
- Pliskova, J., F. Kovarik, O. Kosulic, and F. St'ahlavsky. 2016. Description of a new species of Heterometrus Ehrenberg, 1828 (Scorpiones: Scorpionidae) from Thailand with remarks about the utilization of cytogenetic data in taxonomy of the genus. Annales Zoologici (Warszawa) 66: 467-476.
- Pocock, R.I. 1888. On the African specimens of the genus Scorpio (Linn.), contained in the collection of the British Museum. Annals and Magazine of Natural History (Ser. 6) 2: 245-255.
- Pocock, R.I. 1890. Report upon a small collection of scorpions and centipedes sent from Madras by Mr. Edgar Thurston, of the Government Central Museum. Annals and Magazine of Natural History (Ser. 6) 5: 236-259.
- Pocock, R.I. 1892. Descriptions of two new genera of scorpions, with notes upon some species of Palamnaeus. Annals and Magazine of Natural History (Ser. 6) 9: 38-49.
- Pocock, R.I. 1893. Report upon a small collection of scorpions sent to the British Museum by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Branch of the Royal Asiatic Society 7: 295-312.
- Pocock, R.I. 1894a. A small contribution to our knowledge of the scorpions of India. Annals and Magazine of Natural History (Ser. 6) 13: 72-84.
- Pocock, R.I. 1894b. Supplementary note on the Arachnida and Myriapoda of the Mergui Archipelago: with descriptions of some new species from Siam and Malaysia. Journal of the Linnean Society 24: 316-318.
- Pocock, R.I. 1894c. Scorpions from the Malay Archipelago. In M. Weber (editor), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien 3: 84-99. Leiden: E.J. Brill Verlag.
- Pocock, R.I. 1896a. On a new generic distinction between the scorpions of the genera Scorpio and Palamnaeus. Annals and Magazine of Natural History (Ser. 6) 18: 75-77.
- Pocock, R.I. 1896b. How and why scorpions hiss. Natural Science 9: 17-25.
- Pocock, R.I. 1896c. On the scorpions, centipedes and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History (Ser. 6) 17: 425-444.
- Pocock, R.I. 1897. Descriptions of some new species of scorpions from India. Journal of the Bombay Natural History Society 11: 102-117.
- Pocock, R.I. 1899. Diagnoses of some new Indian Arachnida. Journal of the Bombay Natural History Society 12: 744-753.
- Pocock, R.I. 1900a. Arachnida. In W.T. Blandford (editor), The fauna of British India, including Ceylon and Burma. London: Taylor and Francis, 279 pp.
- Pocock, R.I. 1900b. The scorpions of the genus Heterometrus. Annals and Magazine of Natural History (Ser. 7) 5: 362-365.
- Pocock, R.I. 1902. A contribution to the systematics of scorpions. I. Some corrections in nomenclature. II. Notes on some species of Parabuthus contained in the British Museum. III. Descriptions of some new and old species. Annals and Magazine of Natural History (Ser. 7) 10: 364-380.
- Pointer, M. 1991. The genus Heterometrus and the family Scorpionidae. Journal of the British Tarantula Society 7: 23-25.
- Polis, G.A. 1990. Ecology. In G.A. Polis (editor), The biology of scorpions: 247-340. Stanford, CA: Stanford University Press.
- Polis, G.A., and W.R. Lourenco. 1986. Sociality among scorpions. In J.A. Barrientos (editor), Actas X Congreso de Aracnologia, Jaca (Espana), Septiembre 1986, 1: 111-115. Barcelona: Instituto Pirenaico de Ecologia (C.S.I.C.) Grupo de Aracnologia (Asoc. Esp. Entomol.)
- Polis, G.A., and W.D. Sissom. 1990. Life history. In G.A. Polis (editor), The biology of scorpions: 161-223. Stanford, CA: Stanford University Press.
- Poljansky, I. 1903. Wissenschaftiche Mitteilungen. 1. Zur Embroyologie des Skorpions. Zoologischer Anzeiger 27: 49-58.
- Prendini, L. 2000a. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): an exemplar approach. Cladistics 16: 1-78.
- Prendini, L. 2000b. Chelicerata (Scorpiones). In A.H. Kirk-Spriggs and E. Marais (editors), Daures - Biodiversity of the Brandberg Massif, Namibia. Cimbebasia Memoir 9: 109-120.
- Prendini, L. 2001a. A review of synonyms and subspecies in the genus Opistophthalmus C.L. Koch (Scorpiones: Scorpionidae). African Entomology 9: 17-48.
- Prendini, L. 2001b. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. In V. Fet and P.A. Selden (editors), Scorpions 2001: In memoriam Gary A. Polis: 113-138. Burnham Beeches, Bucks, U.K.: British Arachnological Society.
- Prendini, L. 2003. A new genus and species of bothriurid scorpion from the Brandberg Massif, Namibia, with a reanalysis of bothriurid phylogeny and a discussion of the phylogenetic position of Lisposoma Lawrence. Systematic Entomology 28: 1-24.
- Prendini, L. 2004. On the scorpions of Gabon and neighbouring countries, with a reassessment of the synonyms attributed to Babycurus buettneri Karsch and a redescription of Babycurus melanicus Kovarik. In B.L. Fisher (editor), Monts Doudou, Gabon. A floral and faunal inventory with reference to elevational variation. Memoirs of the California Academy of Sciences 28: 235-267.
- Prendini, L. 2005. Scorpion diversity and distribution in southern Africa: pattern and process. In B.A. Huber, B.J. Sinclair, and K.-H. Lampe (editors), African biodiversity: molecules, organisms, ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn: 25-68. New York: Springer Verlag.
- Prendini, L. 2009. Assembling the scorpion Tree of Life: 'High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)' reanalysed. In J. Arrivillaga, M. El Souki, and B. Herrera, B. (editors), Monografia digital sociedad venezolana de entomologia 2: 1-41. Caracas: Sociedad Venezolana de Entomologia.
- Prendini, L. 2011. Order Scorpiones C.L. Koch, 1850. In Z.-Q. Zhang (editor), Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 115-117.
- Prendini, L. 2016. Redefinition and systematic revision of the East African scorpion genus Pandinoides (Scorpiones: Scorpionidae) with critique of the taxonomy of Pandinus, sensu lato. Bulletin of the American Museum of Natural History 407: 1-66.
- Prendini, L. and T.L. Bird. 2008. Scorpions of the Brandberg Massif, Namibia: species richness inversely correlated with altitude. African Invertebrates 49: 77-107.
- Prendini, L., and W.C. Wheeler. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics 21: 446-494.
- Prendini, L., T.M. Crowe, and W.C. Wheeler. 2003. Systematics and biogeography of the family Scorpionidae Latreille, with a discussion of phylogenetic methods. Invertebrate Systematics 17: 185-259.
- Pringle, J.W.S. 1955. The function of the lyriform organs of arachnids. Journal of Experimental Biology 32: 270-278.
- Purcell, W.F. 1899. On the species of Opisthophthalmus in the collection of the South African Museum, with descriptions of some new forms. Annals of the South African Museum 1: 131-180.
- Rajyalakshmi, A., P. Umamaheshwara Reddy, B. Narasimha Rao, and T. Gopalakrishna Reddy. 1991. The disposition of carbaryl in the hepatopancreas, haemolymph and embryos of the viviparous scorpion, Heterometrus fulvipes (Koch) during gestation. Xenobiotica 21: 429-432.
- Ravindranath, M.H. 1974. The hemocytes of a scorpion Palamnaeus swammerdami. Journal of Morphology 144: 1-10.
- Robert, J.-Y. 1999. Reproduction et croissance en captivite d' Heterometrus spinifer (Scorpiones, Scorpionidae). Arachnides 41: 19-27.
- Roewer, C.F. 1929. Sud-indische Skorpione, Chelonethi und Opilioniden. Revue Suisse de Zoologie 36: 609-613.
- Roewer, C.F. 1943. Uber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg. Senckenbergiana 26: 205-244.
- Ronquist, F., and J.P. Huelsenbeck. 2003. MrBayes 3: Bayesian phylogenetic inference under mixed models. Bioinformatics 19: 1572-1574.
- Root, T.M. 1990. Neurobiology. In G.A. Polis (editor), The biology of scorpions: 341-413. Stanford, CA: Stanford University Press.
- Rosin, R., and A. Shulov. 1961. Sound production in scorpions. Science 133: 1918-1919.
- Rosin, R., and A. Shulov. 1963. Studies of the scorpion Nebo hierichonticus. Proceedings of the Zoological Society of London 140: 547-575.
- Rossi, A. 2014a. Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic and Djibouti (Scorpiones: Scorpionidae). Onychium 10: 10-31.
- Rossi, A. 2014b. The fragmented peri-Saharan distribution of the subgenus Pandinurus Fet, 1997 with the description of a new species from Chad (Scorpiones, Scorpionidae, Pandinus). Serket 14: 6-14.
- Rossi, A. 2014c. New data on the rare species Pandinus nistriae Rossi, 2014 (Scorpiones: Scorpionidae). Arachnides 72: 3-12.
- Rossi, A. 2014d. Clarification of the type locality of Pandinus ulderigoi Rossi, 2014 with notes on the scorpions protected by CITES (Scorpiones: Scorpionidae). Arachnologische Mitteilungen 49: 47-54.
- Rossi, A. 2015a. Sui sottogeneri di Pandinus Thorell, 1876 con revisione del genere Pandinurus Fet, 1997 stat. n. e descrizione di sette nuove specie e tre nuovi sottogeneri (Scorpiones: Scorpionidae). Onychium 11: 10-66.
- Rossi, A. 2015b. Due ulteriori nuove specie del sottogenere Pandiborellius Rossi, 2015 dal Corno d'Africa (Scorpiones, Scorpionidae). Arachnida 1: 2-36.
- Rossi, A. 2015c. Una revision preliminare del genere Pandinus Thorell, 1876 con la descrizione du un nuovo sottogenere e due nuove specie dall'Etiopia (Scorpiones, Scorpionidae). Arachnida 3: 37-52.
- Rossi, A. 2016a. Atlas of scorpions. Volume 1. (Scorpionidae: Pandininae: Pandinoides). Rivista Aracnologica Italiana 8 (Suppl.): 1-148.
- Rossi, A. 2016b. On Pandinus duffmackayi Prendini, 2016, a junior synonym of Pandinoides (Pandinoides) mariachiarae Rossi, 2016, with comments on the recent revisions of the genus Pandinoides Fet, 2000 proposed independently by Rossi and Prendini. Rivista Aracnologica Italiana 10: 45-48.
- Rubio, M. 2000. Scorpions. A complete pet owner's manual. Barrons Educational Series.
- Rutin, J. 1996. The burrowing activity of scorpions (Scorpio maurus palmatus) and their potential contribution to the erosion of hambra soils in Karkur, central Israel. Geomorphology 15: 159-168.
- Santibanez-Lopez, C.E., O.F. Francke, and L. Prendini. 2014a. Phylogeny of the North American scorpion genus Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) based on morphology, nuclear and mitochondrial DNA. Arthropod Systematics and Phylogeny 72: 257-279.
- Santibanez-Lopez, C.E., O.F. Francke, and L. Prendini. 2014b. Kolotl, n. gen. (Scorpiones: Diplocentridae), a new scorpion genus from Mexico. American Museum Novitates 3815: 1-28.
- Sasirababu, K. 1961. "Giant" fibres in the central nervous system of scorpion. Journal of Animal Morphology and Physiology 8: 11-18.
- Sasirababu, K., and S.A.T. Venkatachari. 1966. Certain anatomical features of the ventral nerve cord of the scorpion, Heterometrus fulvipes. Journal of Animal Morphology and Physiology 13: 22-33.
- Scheuring, L. 1912. Uber ein neues Sinnesorgan bei Heterometrus longimanus Hbst. Zoologischer Anzeiger 40: 370-374.
- Schiejok, H. 1997. Einige Gedanken zur Haltung von Pandinus imperator (C.L. Koch, 1842). Skorpion- News 2: 39-43.
- Schultze, W. 1927. Biology of the large Philippine forest scorpion. Philippine Journal of Science 32: 375-388.
- Sergent, E. 1938. Iconographie des scorpions de l'Afrique du Nord. Archives de l'Institut Pasteur d'Algerie 16: 513-522.
- Sergent, E. 1941. Sur le postabdomen (queue) del quelques scorpions de l'Afrique du Nord. Archives de l'Institut Pasteur d'Algerie 19: 353-357.
- Shachak, M., and S. Brand. 1983. The relationship between sit-and-wait foraging strategy and dispersal in the desert scorpion Scorpio maurus palmatus. Oecologia 60: 371-377.
- Sharma, P.P., R. Fernandez, L.A. Esposito, E. Gonzalez- Santillan, and L. Monod. 2015. Phylogenomic resolution of scorpions reveals multilevel discordance with morphological phylogenetic signal. Proceedings of the Royal Society B, Biological Sciences 282: 1-10.
- Shivashankar, T. 1992. The importance of burrowing for the scorpion Heterometrus fulvipes Koch (Arachnida). Journal of Soil Biology and Ecology 12: 134- 138.
- Shivashankar, T. 1994. Advanced subsocial behaviour in the scorpion Heterometrus fulvipes Koch (Arachnida). Journal of Bioscience 19: 81-90.
- Shivashankar, T., and G.K. Veeresh. 1991. Advantages of burrowing by a tropical scorpion, Heterometrus fulvipes (Arachnida: Scorpionidae). In G.K. Veeresh, D. Rajagopal and C.A. Viraktamath (editors), Advances in management and conservation of soil fauna: 697-702. New Delhi: Oxford.
- Shultz, J.W. 2007. Morphology of the prosomal endoskeleton of Scorpiones (Arachnida) and a new hypothesis for the evolution of cuticular cephalic endoskeletons in arthropods. Arthropod Structure and Development 36: 77-102.
- Simard, J.M., and D.D. Watt. 1990. Venoms and toxins. In G.A. Polis (editor), The biology of scorpions: 414-444. Stanford, CA: Stanford University Press.
- Simon, E. 1872a. Arachnides de Syrie rapportes par M. Charles Piochard de la Brulerie (scorpions et Galeodes). Annales de la Societe Entomologique de France 2: 247-264.
- Simon, E. 1872b. Etudes sur les scorpions. Revision des Heterometrus du groupe de l' H. afer, L. Revue et Magasin de Zoologie Pure et Appliquee 23: 51- 59; 97-101.
- Simon, E. 1879. 3e Ordre. Scorpiones. In Les arachnides de France. VII. Contenant les ordres des Chernetes, Scorpiones et Opiliones: 79-115. Paris: Roret.
- Simon, E. 1880. [Scorpions du Mossoul (ancienne Ninive), sur le Tigre, en Mesopotamie.] Bulletin de la Societe Entomologique de France 10: 29.
- Simon, E. 1884a. Arachnides recueillis en Birmanie par M. le Chevalier J.B. Comotto et appartenent au Musee Civique d'Histoire Naturelle de Genes. Annali del Museo Civico di Storia Naturale di Genova 20: 37-48 (361-372).
- Simon, E. 1884b. Etudes arachnologiques. 15e Memoire. XXII. Arachnides recueillis par M. l'abbe A. David a Smyrne, a Beirout et a Akbes en 1883. Annales de la Societe Entomologique de France 4: 181-196.
- Simon, E. 1885. Materiaux pour servir a la faune arachnologique de l'Asie meridionale. Annales de la Societe Entomologique de France 7: 22-39.
- Simon, E. 1887. Etude sur les arachnides de l'Asie meridionale faisant partie des collections de l'Indian Museum (Calcutta). I. Arachnides recueillis a Tavoy (Tenasserim) par Moti Ram. Journal of the Asiatic Society of Bengal 56: 101-117.
- Simon, E. 1892. Liste des arachnides recueillis en Syrie par M. le docteur Theod. Barrois. Revue Biologique du Nord de la France 5: 80-84.
- Simon, E. 1893. Arachnides de l'Archipel Malais recueillis par MM. M. Bedot et C. Pictet. Revue Suisse de Zoologie 1: 319-328.
- Simon, E. 1899. Contributions a la faune de Sumatra. Arachnides recueillis par M.J.L. Weyers, a Sumatra (2e memoire). Annales de la Societe Entomologique de Belgique 43: 78-125.
- Simon, E. 1901. On the arachnids collected during the "Skeat Expedition" to the Malay Peninsula, 1899- 1900. Proceedings of the Zoological Society of London 2: 45-84.
- Simon, E. 1904. Arachnides recueillis par M.A. Pavie en Indo-Chine. In Mission Pavie en Indo-Chine 1879- 1895. Etudes diverses III. Recherches sur l'histoire naturelle de l'Indochine orientale: 270-295. Paris: Ernest Leroux.
- Simon, E. 1905. Voyage de M. Maurice Maindron dans l'Inde meridionale. Arachnides. Annales de la Societe Entomologique de France 74: 160-180.
- Simon, E. 1910. Revision des scorpions d'Egypte. Bulletin de la Societe Entomologique d'Egypte (Cairo) 1910: 57-87.
- Sinha, R.C. 1966. Effect of starvation on the concentration of ascorbic acid in the pedipalp muscle of the scorpion (P. bengalensis). Experientia 22: 221- 222.
- Sinha, R.C., and M.S. Kanungo. 1967. Effects of starvation on the scorpion Palamnaeus bengalensis. Physiological Zoology 40: 386-390.
- Sissom, W.D. 1990. Systematics, biogeography and paleontology. In G.A. Polis (editor), The biology of scorpions: 64-160. Stanford, CA: Stanford University Press.
- Skaife, S.H. 1920. Animal life in South Africa. Cape Town: Maskew Miller.
- Skaife, S.H. 1954. South African nature notes. 2nd ed. Cape Town: Maskew Miller.
- Soleglad, M.E., and V. Fet. 2003a. High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni). Euscorpius 11: ii+1-175.
- Soleglad, M.E., and V. Fet. 2003b. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius 5: 1-34.
- Soleglad, M.E., V. Fet, and F. Kovarik. 2005. The systematic position of the scorpion genera Heteroscorpion Birula, 1903 and Urodacus Peters, 1861 (Scorpiones: Scorpionoidea). Euscorpius 20: 1-38.
- Sreenivasa-Reddy, R.P. 1968. Contribution a la connaissance des scorpions de l'Inde. 5. Le genre Iomachus Pocock, 1893 (Scorpionidae, Ischnurinae). Bulletin du Museum National d'Histoire Naturelle, Paris 40: 759-767.
- Srivastava, D.S. 1955. Maxillary processes and mechanism of feeding in scorpions. Sugar University Journal 2: 85-91.
- Srivastava, D.S., and M.S. Kanungo. 1966. Metabolism of the liver of the scorpion, Palamnaeus bengalensis. Comparative Biochemistry and Physiology 19: 629-632.
- Stahnke, H.L. 1974. An estimate of the number of taxa in the order Scorpionida. BioScience 24: 339.
- Stamatakis, A. 2006. RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models. Bioinformatics 22: 2688- 2690.
- Stamatakis, A. 2014. RAxML version 8: a tool for phylogenetic analysis and post-analysis of large phylogenies. Bioinformatics 30: 1312-1313.
- Stockmann, R. 1979. Developpement postembrynonaire et cycle d'intermue chez un scorpion Buthidae - Buthotus minax occidentalis. Bulletin du Museum National d'Histoire Naturelle, Paris 4: 405-420.
- Stockwell, S.A. 1989. Revision of the phylogeny and high classification of scorpions (Chelicerata). Ph.D. dissertation, University of California, Berkeley.
- Subburam, V., and T. Gopalakrishna Reddy. 1978a. Breeding season and gestation period of the scorpion Heterometrus fulvipes. Journal of the Bombay Natural History Society 75: 513-514.
- Subburam, V., and T. Gopalakrishna Reddy. 1978b. Changes of glucose and glycogen content in the maternal and embryonic tissues of the viviparous scorpion Heterometrus fulvipes during gestation period. Current Science 47: 665-667.
- Subburam, V., and T. Gopalakrishna Reddy. 1979. Changes in the protein content of the maternal and embryonic tissues of the viviparous scorpion Heterometrus fulvipes during gestation period. Current Science 48: 853-856.
- Subburam, V., and T. Gopalakrishna Reddy. 1980a. Changes in the lipid content of the maternal and embryonic tissues and variations in the maternal hepatosomatic index during the gestation period of the viviparous scorpion, Heterometrus fulvipes. Current Science 49: 658-661.
- Subburam, V., and T. Gopalakrishna Reddy. 1980b. Changes in the oxygen consumption of the maternal animal, and maternal and embryonic tissues during the gestation period of the viviparous scorpion, Heterometrus fulvipes. Current Science 49: 138-140.
- Subburam, V., and T. Gopalakrishna Reddy. 1981. Morphology of the embryonic stages of the viviparous scorpion, Heterometrus fulvipes: a photographic study. Journal of Morphology 169: 275-281.
- Subburam, V., and T. Gopalakrishna Reddy. 1989. Functional morphology of the appendix of the viviparous scorpion, Heterometrus fulvipes (Koch) (Arachnida: Scorpionidae). International Journal of Insect Morphology and Embryology 18: 329-333.
- Sundevall, C.J. 1833. Conspectus arachnidum. Londini Gothorum: C.F. Berling. 39 pp.
- Sureshan, P.M., D.B. Bastawade, and C. Radhakrishnan. 2007a. Taxonomic studies on a collection of scorpions (Scorpiones: Arachnida) from Western Ghats in Kerala, India with two new distribution records. Zoos' Print Journal 22: 2903-2908.
- Sureshan, P.M., D.B. Bastawade, and C. Radhakrishnan. 2007b. Scorpion fauna (Arachnida: Scorpionida) of Parambikulam Wild Life Sanctuary, Kerala, India. Records of the Zoological Survey of India 107: 49-54.
- Tahir, H.M., and L. Prendini. 2014. Redescription of Heterometrus latimanus and confirmation of the genus Heterometrus (Scorpiones: Scorpionidae) in Pakistan. American Museum Novitates 3805: 1-23.
- Takashima, H. 1941. [Scorpiones and Pedipalpi from the islands of Riu-Kiu.] Biogeographica (Tokyo) 3: 273-285. [in Japanese]
- Takashima, H. 1945. [Scorpions of eastern Asia.] Acta Arachnologica 9: 68-106. [in Japanese]
- Takashima, H. 1952. [Miscellaneous notes on scorpions.] Acta Arachnologica 13: 26-36. [in Japanese]
- Talal, S., et al. 2015. Scorpion speciation in the Holy Land: multilocus phylogeography corroborates diagnostic differences in morphology and burrowing behavior among Scorpio subspecies and justifies recognition as phylogenetic, ecological and biological species. Molecular Phylogenetics and Evolution 91: 226-237.
- Tare, T.G., N.E. Vad, and D.M. Renapurkar. 1993. Burrow patterns of the scorpion Heterometrus indus. Medical and Veterinary Entomology 7: 102-104.
- Thorell, T. 1876a. On the classification of scorpions. Annals and Magazine of Natural History (Ser. 4) 17: 1-15.
- Thorell, T. 1876b. Etudes Scorpiologiques. Atti della Societa Italiana di Scienze Naturali 19: 75-272.
- Thorell, T. 1888. Pedipalpi e scorpioni dell'Archipelago Malesi conservati nel Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova "Giacomo Doria" 26: 327-428.
- Thorell, T. 1890. Aracnidi di Nias e di Sumatra raccolti nel 1886 dal Sign. E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova "Giacomo Doria" 30: 5-106.
- Thorell, T. 1893. Scorpiones exotici R. Musei Historiae Naturalis Florentini. Bolletino della Societa Entomologica Italiana 25: 356-387.
- Thorell, T., and G. Lindstrom. 1885. On a Silurian scorpion from Gotland. Bihang till Kungliga Svenska Vetenskapsakademiens Handlingar 21: 1-33.
- Thulsi Rao, K., D.B. Bastawade, M. Sudhakar, S.M.M. Javed, and I.S.R. Krishna. 2005. Arachnid fauna of Nallamalai Region Eastern Ghats, Andhra Pradesh, India. Records of the Zoological Survey of India, Occasional Paper 239: 1-42.
- Tikader, B.K. 1973. Arachnida of the Deccan area. Bulletin of the Indian National Science Academy 45: 260-276.
- Tikader, B.K. 1987. The scorpions. In Handbook Indian spiders: 20-38. Calcutta: Zoological Survey of India.
- Tikader, B.K., and D.B. Bastawade. 1983. Fauna of India, vol. 3. Scorpions (Scorpionida: Arachnida). Zoological Survey of India. Pune: Sangam Press. 671 pp.
- Toye, S.A. 1970. Some aspects of the biology of two common species of Nigerian scorpions. Journal of Zoology, London 162: 1-9.
- Uthaman, M., and Y. Srinivasa Reddy. 1979. Bimodal oscillation in the oxygen consumption of the isolated tissues of the scorpion Heterometrus fulvipes. Journal of Arid Environments 2: 347-353.
- Vachon, M. 1950. Etudes sur les scorpions. III (suite). Description des scorpions du nord de l'Afrique. Archives de l'Institut Pasteur d'Algerie 28: 152-216.
- Vachon, M. 1952a. Etudes sur les scorpions. Alger: Institut Pasteur d'Algerie, 482 pp.
- Vachon, M. 1952b. La Reserve Naturelle Integrale du Mt. Nimba I. Scorpions. Mission M. Lamotte en Guinee (1942). Memoires de l'Institut Francais de Afrique Noire 19: 9-15.
- Vachon, M. 1953. Quelques remarques sur les frontieres en biogeographie a propos de la Ligne Wallace et des scorpions indonesiens. Comptes Rendus des Seances de la Societe de Biogeographie 30: 6-12.
- Vachon, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris 35: 161-166.
- Vachon, M. 1974 ("1973"). Etude des caracteres utilises pour classer les familles et les genres de scorpions (arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris 140: 857-958.
- Vachon, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d'Histoire naturelle de Geneve III.). Revue Suisse de Zoologie 89: 77-114.
- Vachon, M., and T. Abe. 1988. Colonization of the Krakatau Islands (Indonesia) by scorpions. Acta Arachnologica 37: 23-33.
- Vachon, M., and W.R. Lourenco. 1985. Scorpions cavernicoles du Sarawak (Borneo). Chaerilus chapmani n. sp. (Chaerilidae) et Lychas hosei (Pocock, 1890) (Buthidae). Memoires Biospeologiques 12: 9-18.
- Vachon, M., B. Dumortier, and R.-G. Busnel. 1958. Enregistrement de stridulations d'un scorpion sudafricain. Bulletin de la Societe Zoologique de France 83: 253-254.
- Vachon, M., R.-G. Busnel, and B. Dumortier. 1960. Production de sons par un scorpion sud-africain (Opisthophthalmus latimanus). Bulletin de la Societe Zoologique de France 85: 79.
- Vachon, M., R. Roy, and M. Condamin. 1970. Le developpement post-embryonnaire du scorpion Pandinus gambiensis Pocock. Bulletin de l'Institut Francais de Afrique Noire (A) 32: 414-432.
- Van Zinderen Bakker, E.M. 1978. Late-Mesozoic and Tertiary palaeoenvironments of the Sahara region. In E.M. van Zinderen Bakker (editor), Antarctic glacial history and world palaeoenvironments: 129- 135. Rotterdam: A.A. Balkema.
- Van Zinderen Bakker, E.M. 1980. Comparison of late- Quaternary climatic evolutions in the Sahara and the Namib-Kalahari region. Palaeoecology of Africa 12: 381-394.
- Venkatachari, S.A.T., and P. Muralikrishna Dass. 1968. Choline esterase activity rhythym in the ventral nerve cord of scorpion. Life Sciences 7: 617-621.
- Venkateswara Rao, P., and S. Govindappa. 1967. Dehyrdogenase activity and its diurnal variations in different muscles of the scorpion Heterometrus fulvipes. Proceedings of the Indian Academy of Sciences, B 66: 243-249.
- Veronika, K., K. Akilan, T. Eswaramohan, and A. Murugananthan. 2013. Diversity and identification key to the species of scorpions (Scorpiones: Arachnida) from Jaffna Peninsula, Sri Lanka. Journal of Entomology and Zoology Studies 1: 70-77.
- Vijayalakshmi, N.R., and P.A. Kurup. 1976. Metabolism of glycosaminoglycans in Heterometrus scaber. Indian Journal of Experimental Biology 14: 10-13.
- Volschenk, E.S., and L. Prendini. 2008. Aops oncodactylus gen. et sp. nov., the first troglobitic urodacid (Chelicerata: Scorpiones), with a re-assessment of cavernicolous, troglobitic and troglomorphic scorpions. Invertebrate Systematics 22: 235-257.
- Volschenk, E.S., C.I. Mattoni, and L. Prendini. 2008. Comparative anatomy of the mesosomal organs of scorpions (Chelicerata, Scorpiones), with implications for the phylogeny of the order. Zoological Journal of the Linnean Society 154: 651-675.
- Vyas, A.B. 1971. Adaptive diversification of the prosomatic appendanges in the scorpion Heterometrus fulvipes. Annals of Zoology 7: 65-80.
- Vyas, A.B. 1974. The cheliceral muscles of the scorpion Heterometrus fulvipes. Bulletin of the Southern California Academy of Sciences 73: 9-14.
- Vyas, A.B., and S.M. Laliwala. 1972. Certain noteworthy features of the circulatory system of Heterometrus fulvipes. Proceedings of the National Academy of Sciences (India) Sect. B, 42: 267-271.
- Wallin, L. 1994. Uppsala University Zoological Museum. Catalog of type specimens (Version 4). 4. Linnean specimens. Uppsala. [Scorpiones: p. 13.]
- Wang, K., et al. 2019. Vibrational receptor of scorpion (Heterometrus petersii): the basitarsal compound slit sensilla. Journal of Bionic Engineering 16: 76-87.
- Warburg, M.R. 2010. Reproductive system of female scorpion: a partial review. Anatomical Record 293: 1738-1754.
- Warburg, M.R. 2012a. Reviewing the structure and function of the scorpion's hepatopancreas. Arthropods 1: 79-93.
- Warburg, M.R. 2012b. Ovulation, fertilization and embryonic development in scorpions: a partial review. African Entomology 20: 60-68.
- Warburg, M.R. 2013. The circadian rhythm and visual elements in scorpions: a review. Arthropods 2: 150- 158.
- Warburg, M.R., and G.A. Polis. 1990. Behavioral responses, rhythms and activity patterns. In G.A. Polis (editor), The biology of scorpions: 224-246. Stanford, CA: Stanford University Press.