Published September 21, 2016 | Version v1
Journal article Open

Redefinition And Systematic Revision Of The East African Scorpion Genus Pandinoides (Scorpiones: Scorpionidae) With Critique Of The Taxonomy Of Pandinus, Sensu Lato

  • 1. Scorpion Systematics Research Group, Division of Invertebrate Zoology, American Museum of Natural History

Description

Prendini, Lorenzo (2016): Redefinition And Systematic Revision Of The East African Scorpion Genus Pandinoides (Scorpiones: Scorpionidae) With Critique Of The Taxonomy Of Pandinus, Sensu Lato. Bulletin of the American Museum of Natural History 2016 (407): 1-66, DOI: 10.1206/0003-0090-407.1.1, URL: http://www.bioone.org/doi/10.1206/0003-0090-407.1.1

Files

source.pdf

Files (12.4 MB)

Name Size Download all
md5:2fb59cfb89593298bf1980f972ebbc08
12.4 MB Preview Download

Linked records

Additional details

Identifiers

References

  • Becker, L. 1880. Etudes sur les scorpions. Annales de la Societe Entomologique de Belgique 24: 134-145.
  • Belfield, W. 1956. A preliminary check list of the West African scorpions and key for their identification. Journal of the West African Science Association 2: 41-47.
  • Birula, A.A. 1913. Arachnologische Beitrage II-IV. (II. Ueber einige Scorpiops -Arten von dem Sudabhange des Himalaya. III. Ueber Pandinus (Pandinops) peelei Poc. und seine Verwandten. IV. Ueber das Vorkommen der gemeinen Perlmutterzecke (Dermacentor reticulatus [Fabr.] - Ixodidae) in den mittleren Teilen Westrusslands). Revue Russe d'Entomologie 13: 416-423.
  • Birula, A.A. 1915a. Arachnologische Beitrage. V. Weitere Bemerkungen uber die Skorpionen-Fauna Britisch Ost-Afrikas. Revue Russe d'Entomologie 15: 50-65.
  • Birula, A.A. 1915b. A general list of the scorpions of British East Africa. Scientific results of the zoological expedition to British East Africa and Uganda made by Prof. V. Dogiel and I. Sokolow 1: 1-31.
  • Birula, A.A. 1927. Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. XXV. Skorpione. Denkschriften der Akademie der Wissenschaften in Wien, Mathemathisch-naturwissenschaftliche Klasse 101: 79-88.
  • Borelli, A. 1901. Materiali per la conoscenza della fauna Eritrea raccolti dal Dott. Paolo Magretti. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Universita di Torino 16: 1-5.
  • Borelli, A. 1919. Missione per la frontiera italo etiopica sotto il comando del Capitano Carlo Citerni. Risulti Zoologici. Scorpioni. Annali del Museo Civico di Storia Naturale di Genova (3) 8: 359-381.
  • Borelli, A. 1925a. Scorpioni nuovi o poco noti della Somalia italiana. Annali del Museo Civico di Storia Naturale di Genova 52: 9-16.
  • Borelli, A. 1925b. Di alcuni scorpioni della Somalia italiana. Annali del Museo Civico di Storia Naturale di Genova 51: 317-326.
  • Caporiacco, L. di. 1937. Su alcuni scorpioni dell'Africa orientale italiana del Civico Museo di Milano. Atti della Societa Italiana di Scienze Naturali 76: 355- 362.
  • Donaldson Smith, A. 1896. Expedition through Somaliland to Lake Rudolf. Geographical Journal 8: 120-137.
  • Donaldson Smith, A. 1897. Through unknown African countries; the first expedition from Somaliland to Lake Lamu. London: Edward Arnold.
  • El-Hennawy, H.K. 1992. A catalogue of the scorpions described from the Arab countries (1758-1990) (Arachnida: Scorpionida). Serket 2: 95-153.
  • Fet, V. 1997. Notes on the taxonomy of some Old World scorpions (Scorpiones: Buthidae, Chactidae, Ischnuridae, Scorpionidae). Journal of Arachnology 25: 245-250.
  • Fet, V. 2000. Family Scorpionidae Latreille, 1802. In V. Fet, W.D. Sissom, G. Lowe, and M.E. Braunwalder, Catalog of the scorpions of the world (1758-1998): 427-486. New York: New York Entomological Society.
  • Fet V., M.E. Soleglad, D.P.A. Neff, and I. Stathi. 2004. Tarsal armature in the superfamily Iuroidea (Scorpiones: Iurida). Revista Iberica de Aracnologia 10: 17-40.
  • Francke, O.F. 1985. Conspectus genericus scorpionorum 1758-1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University 98: 1-32.
  • Francke, O.F., and L. Prendini. 2008. Phylogeny and classification of the giant hairy scorpions, Hadrurus Thorell (Iuridae Thorell): a reappraisal. Systematics and Biodiversity 6: 205-223.
  • Hirst, S. 1911. Scorpions and solifugae collected by Captain S.S. Flower in the Anglo-Egyptian Sudan. Annals and Magazine of Natural History 8: 217- 222.
  • Karsch, F. 1879. Scorpionologische Beitrage. Part I. Mitteilungen des Munchener Entomologischen Vereins 3: 6-22.
  • Koch, C.L. 1841. Die Arachniden. Nurnberg: C.H. Zeh'sche Buchhandlung 9: 1-11.
  • Koch, L. 1875. Aegyptische und abyssinische Arachniden gesammelt von Herrn C. Jickeli beschrieben und abgebildet. Nurnberg: Bauer & Raspe. 96 pp.
  • Kovarik, F. 1992. A check list of scorpions (Arachnida: Scorpiones) in the collections of the zoological department, National Museum in Prague. Acta Societatis Zoologicae Bohemoslovenicae 56: 181-186.
  • Kovarik, F. 1997. A check-list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest. Annales Historico-Naturales Musei Nationalis Hungarici 89: 177-185.
  • Kovarik, F. 1998. Stiri [Scorpions]. Jihlava, Czech Republic: Madagaskar, 176 pp. [in Czech].
  • Kovarik, F. 2000. Pandinus (Pandinops) pococki sp. n. from Somalia, and Pandinus pugilator, a junior synonym of Pandinus (Pandinops) bellicosus comb. n. (Scorpiones, Scorpionidae). Serket 7: 1-7.
  • Kovarik, F. 2002. A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg. Serket 8: 1-23.
  • Kovarik, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones), with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae 67: 133-159.
  • Kovarik, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp.
  • Kovarik, F. 2011. A review of the subgenus Pandinus Thorell, 1876 with descriptions of two new species from Uganda and Ethiopia (Scorpiones: Scorpionidae). Euscorpius 129: 1-18.
  • Kovarik, F. 2012. Review of the subgenus Pandinurus Fet, 1997 with descriptions of three new species (Scorpiones, Scorpionidae, Pandinus). Euscorpius 141: 1-22.
  • Kovarik, F. 2013a. Family Buthidae. In F. Kovarik and A.A. Ojanguren Affilastro (editors), Illustrated catalogue of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas and Sassanidotus: 145-212. Prague: Clarion Production.
  • Kovarik, F. 2013b. Pandinus (Pandinus) trailini sp. n. from Ethiopia (Scorpiones: Scorpionidae), with data on localities and life strategy. Euscorpius 163: 1-14.
  • Kovarik, F., and S. Whitman. 2005. Cataloghi del Museo di Storia Naturale dell'Universita di Firenze - sezione di zoologia "La Specola" XXII. Arachnida Scorpiones. Tipi. Addenda (1998-2004) e checklist della collezione (Euscorpiinae esclusi). Atti della Societa Toscana di Scienze Naturali, Memorie 111: 103-119.
  • Kovarik, F., G. Lowe, D. Hoferek, J. Pliskova, and F. Stahlavsky. 2016. Scorpions of Ethiopia. Part IV. Genus Uroplectes Peters, 1861 (Scorpiones: Buthidae). Euscorpius 217: 1-14.
  • Kraepelin, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (editor), Das Tierreich 8 (Arachnoidea): 1-265. Herausgegeben von der Deutschen Zoologischen Gesellschaft. Berlin: R. Friedlander und Sohn Verlag.
  • Kraepelin, K. 1901. Catalogue des scorpions des collections du Museum d'Histoire Naturelle de Paris. Bulletin du Museum National d'Histoire Naturelle, Paris 7: 265-273.
  • Kraepelin, K. 1908. Die sekundaren Geschlechtscharaktere der Skorpione, Pedipalpen und Solifugen. Mitteilungen aus dem Naturhistorischen Museum (2. Beiheft zum Jahrbuch der Hamburgischen wissenschaftlichen Anstalten) 25: 181-225.
  • Kraepelin, K. 1913. Neue Beitrage zur Systematik der Gliederspinnen. III. A. Bemerkungen zur Skorpionenfauna Indiens. B. Die Skorpione, Pedipalpen und Solifugen Deutsch-Ost-Afrikas. Mitteilungen aus dem Naturhistorischen Museum (2. Beiheft zum Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 1912) 30: 123-196.
  • Lamoral, B.H. 1979. The scorpions of Namibia (Arachnida: Scorpionida). Annals of the Natal Museum 23: 497-784.
  • Lamoral, B.H., and S.C. Reynders. 1975. A catalogue of the scorpions described from the Ethiopian faunal region up to December 1973. Annals of the Natal Museum 22: 489-576.
  • Lourenco, W.R. 1987. Revision systematique des scorpions du genre Opisthacanthus (Scorpiones, Ischnuridae). Bulletin du Museum National d'Histoire Naturelle, Paris (4) 9: 887-931.
  • Lourenco, W.R. 2009. Reanalysis of the genus Scorpio Linnaeus 1758 in sub-Saharan Africa and description of one new species from Cameroon (Scorpiones, Scorpionidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 15: 99-113.
  • Lourenco, W.R. 2014. Further considerations on the identity and distribution of Pandinus imperator (C.L. Koch, 1841) and description of a new species from Cameroon (Scorpiones: Scorpionidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 17: 139-151.
  • Lourenco, W.R., and J.L. Cloudsley-Thompson. 1996. Recognition and distribution of the scorpions of the genus Pandinus Thorell, 1876 accorded protection by the Washington Convention. Biogeographica, Paris 72: 133-143.
  • Loveridge, A. 1925. Notes on East African scorpions and Solifugae, collected 1916-23. Proceedings of the Zoological Society of London 1925: 303-309.
  • Moriggi, M. 1941. Gli scorpioni dell'Africa orientale italiana. Rivista di Biologia Coloniale 4: 77-103.
  • Pocock, R.I. 1888. On the African specimens of the genus Scorpio (Linn.), contained in the collection of the British Museum. Annals and Magazine of Natural History (6) 2: 245-255.
  • Pocock, R.I. 1890. Descriptions of two new species of scorpions brought by Emin Pasha from the inland parts of East Africa. Annals and Magazine of Natural History (6) 6: 98-101.
  • Pocock, R.I. 1896. On the scorpions, centipedes and millipedes obtained by Dr. Gregory on his expedition to Mount Kenia, East Africa. Annals and Magazine of Natural History (6) 17: 425-444.
  • Pocock, R.I. 1897a. Descriptions of two new species of scorpions from East Africa. Annals and Magazine of Natural History (6) 19: 116-119.
  • Pocock, R.I. 1897b. Solifugae, Scorpiones, Chilopoda and Diplopoda (Appendix C). In A. Donaldson Smith, Through unknown African countries; the first expedition from Somaliland to Lake Lamu: 392-407. London: Edward Arnold.
  • Pocock, R.I. 1898a. The Arachnida from the regions of Lake Nyassa and Tanganyika contained in the collection of the British Museum. Annals and Magazine of Natural History (7) 2: 429-448.
  • Pocock, R.I. 1898b. On the scorpions, spiders and solpugas, collected by Mr C. Stewart Betton in British East Africa. Proceedings of the Zoological Society of London 1898: 497-524.
  • Pocock, R.I. 1899. On the scorpions, pedipalps and spiders from Tropical West Africa represented in the collection of the British Museum. Proceedings of the Zoological Society of London 1899: 833- 885.
  • Pocock, R.I. 1900a. On a collection of insects and arachnids made in 1895 and 1897, by Mr C.A.V. Peel, F.Z.S., in Somaliland, with descriptions of new species. 9. The Chilopoda and Arachnida. Proceedings of the Zoological Society of London 1900: 48-55.
  • Pocock, R.I. 1900b. On a collection of insects and arachnids made in 1895 and 1897, by Mr C.V.A. Peel, F.Z.S., in Somaliland, with descriptions of new species. 10. General list of the scorpions of Somalil- and and the Boran Country. Proceedings of the Zoological Society of London 1900: 55-63.
  • Pocock, R.I. 1902. A contribution to the systematics of scorpions. I. Some corrections in nomenclature. II. Notes on some species of Parabuthus contained in the British Museum. III. Descriptions of some new and old species. Annals and Magazine of Natural History (7) 10: 364-380.
  • Prendini, L. 2000a. Chelicerata (Scorpiones). In A.H. Kirk-Spriggs and E. Marais (editors), Daures - Biodiversity of the Brandberg Massif, Namibia. Cimbebasia Memoir 9: 109-120.
  • Prendini, L. 2000b. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): an exemplar approach. Cladistics 16: 1-78.
  • Prendini, L. 2001a. A review of synonyms and subspecies in the genus Opistophthalmus C.L. Koch (Scorpiones: Scorpionidae). African Entomology 9: 17-48.
  • Prendini, L. 2001b. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. In V. Fet and P.A. Selden (editors), Scorpions 2001. In memoriam Gary A. Polis: 113-138. Burnham Beeches, Buckinghamshire, UK: British Arachnological Society.
  • Prendini, L. 2001c. Two new species of Hadogenes (Scorpiones, Ischnuridae) from South Africa, with a redescription of Hadogenes bicolor and a discussion on the phylogenetic position of Hadogenes. Journal of Arachnology 29: 146-172.
  • Prendini, L. 2004. On the scorpions of Gabon and neighbouring countries, with a reassessment of the synonyms attributed to Babycurus buettneri Karsch and a redescription of Babycurus melanicus Kovarik. In B.L. Fisher (editor), Monts Doudou, Gabon. A floral and faunal inventory with reference to elevational variation. Memoirs of the California Academy of Sciences 28: 235-267.
  • Prendini, L. 2005. Scorpion diversity and distribution in southern Africa: pattern and process. In B.A. Huber, B.J. Sinclair, and K.-H. Lampe (editors), African biodiversity: molecules, organisms, ecosystems. Proceedings of the 5th International Symposium on Tropical Biology, Museum Alexander Koenig, Bonn: 25-68. New York: Springer Verlag.
  • Prendini, L. 2006. New South African flat rock scorpions (Liochelidae: Hadogenes). American Museum Novitates 3502: 1-32.
  • Prendini, L., and W.C. Wheeler. 2005. Scorpion higher phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics 21: 446-494.
  • Prendini, L., T.M. Crowe, and W.C. Wheeler. 2003. Systematics and biogeography of the family Scorpionidae Latreille, with a discussion of phylogenetic methods. Invertebrate Systematics 17: 185-259.
  • Probst, P.J. 1973. A review of the scorpions of East Africa with special regard to Kenya and Tanzania. Acta Tropica 30: 312-335.
  • Roewer, C.F. 1943. Uber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg. Senckenbergiana 26: 205-244.
  • Rossi, A. 2014a. Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic and Djibouti (Scorpiones: Scorpionidae). Onychium 10: 10-31.
  • Rossi, A. 2014b. The fragmented peri-Saharan distribution of the subgenus Pandinurus Fet, 1997 with the description of a new species from Chad (Scorpiones, Scorpionidae, Pandinus). Serket 14: 6-14.
  • Rossi, A. 2014c. New data on the rare species Pandinus nistriae Rossi, 2014 (Scorpiones: Scorpionidae). Arachnides 72: 3-12.
  • Rossi, A. 2014d. Clarification of the type locality of Pandinus ulderigoi Rossi, 2014 with notes on the scorpions protected by CITES (Scorpiones: Scorpionidae). Arachnologische Mitteilungen 49: 47-54.
  • Rossi, A. 2015a. Sui sottogeneri di Pandinus Thorell, 1876 con revisione del genere Pandinurus Fet, 1997 stat. n. e descrizione di sette nuove specie e tre nuovi sottogeneri (Scorpiones: Scorpionidae). Onychium 11: 10-66.
  • Rossi, A. 2015b. Due ulteriori nuove specie del sottogenere Pandiborellius Rossi, 2015 dal Corno d'Africa (Scorpiones, Scorpionidae). Arachnida 1: 2-36.
  • Rossi, A. 2015c. Una revision preliminare del genere Pandinus Thorell, 1876 con la descrizione du un nuovo sottogenere e due nuove specie dall'Etiopia (Scorpiones, Scorpionidae). Arachnida 3: 37-52.
  • Santibanez-Lopez, C.E., O.F. Francke, and L. Prendini. 2014. Kolotl, n. gen. (Scorpiones: Diplocentridae), a new scorpion genus from Mexico. American Museum Novitates 3815: 1-28.
  • SchiOtz, A. 2003. Alexander Duff-Mackay: in memoriam. 30 December 1939-6 March 2003. Journal of East African Natural History 92: 1-2.
  • Simon, E. 1872. Etudes sur les scorpions. Revue et Magasin de Zoologie Pure et Appliquee (2) 23: 51-59, 97-101.
  • Soleglad, M.E., V. Fet, and M. Graham. 2014. Kovarikia, a new scorpion genus from California, USA (Scorpiones: Vaejovidae). Euscorpius 185: 1-22.
  • Spawls, S., K. Howell, R. Drewes, and J. Ashe. 2004. A field-guide to the reptiles of East Africa. London: A & C Black, 543 pp.
  • Tahir, H.M., and L. Prendini. 2014. Redescription of Heterometrus latimanus and confirmation of the genus Heterometrus (Scorpiones: Scorpionidae) in Pakistan. American Museum Novitates 3805: 1-23.
  • Thorell, T. 1876a. On the classification of scorpions. Annals and Magazine of Natural History 4: 1-15.
  • Thorell, T. 1876b. Etudes scorpiologiques. Atti della Societa Italiana di Scienze Naturali 19: 75-272.
  • Thorell, T. 1893. Scorpiones exotici R. Musei Historiae Naturalis Florentini. Bollettino della Societa Entomologica Italiana 25: 356-387.
  • Vachon, M. 1967. Le grand scorpion du Senegal: Pandinus gambiensis Pocock, 1899 doit etre considere comme une veritable espece et non comme une sous-espece de Pandinus imperator C.L. Koch, 1842. Bulletin de l'Institut Fondamental d'Afrique Noire (A) Sciences Naturelles 29: 1534-1537.
  • Vachon, M. 1973 [1974]. Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie, Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris (3) 140: 857-958.
  • Werner, F. 1916. Ueber einige Skorpione und Gliederspinnen des Naturhistorischen Museums in Wiesbaden. Jahrbucher des Nassauischen Vereins fur Naturkunde 69: 79-97.
  • Werner, F. 1934. Scorpiones, Pedipalpi. In H.G. Bronns Klassen und Ordnungen des Tierreichs. Akademische Verlaggesellschaft, Leipzig. 5, IV, 8, Lief. 1-2 (Scorpiones): 1-316.
  • Werner, F. 1936. Neu-Eingange von Skorpionen im Zoologischen Museum in Hamburg. Festschrift zum 60. Geburtstage von Professor Dr. Embrik Strand 2: 171-193.