Published December 31, 2017
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FIGURES 47–54. 47, 49, 51, 53 in Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)
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FIGURES 47–54. 47, 49, 51, 53. Male head, dorsal view; 48, 50, 52, 54. Male head, frontal view. 47–48. Digitonthophagus fimator, holotype; 49–50. D. lusinganus, lectotype; 51–52. D. ulcerosus, holotype; 53–54. D. namaquensis, holotype.
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