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Published August 6, 2025 | Version v1
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A new species of the Gonyosoma prasinum species complex (Squamata: Serpentes: Colubridae) from Indochina

Creators

  • 1. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- & Moscow 119991, Russia & Joint Vietnam-Russia Tropical Science and Technology Research Center, Nghia Do, Hanoi, Vietnam
  • 2. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- & Moscow 119991, Russia
  • 3. Sustainable Development Education Center, Tu Liem, Hanoi, Vietnam
  • 4. Joint Vietnam-Russia Tropical Science and Technology Research Center, Nghia Do, Hanoi, Vietnam
  • 5. Institut de Systématique, Évolution et Biodiversité (ISYEB), Muséum National d'Histoire Naturelle, Sorbonne Université, École Pratique des Hautes Études, Université des Antilles, CNRS, CP 30, 57 rue Cuvier, F-75005 Paris, France
  • 6. Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam & College of Medicine and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam *Corresponding authors

Description

Poyarkov, Nikolay A., Bragin, Andrey M., Idiiatullina, Sabira S., Tran, Tuan Anh, Le, Dac Xuan, David, Patrick, Nguyen, Tan Van (2025): A new species of the Gonyosoma prasinum species complex (Squamata: Serpentes: Colubridae) from Indochina. Zootaxa 5673 (3): 355-384, DOI: 10.11646/zootaxa.5673.3.2, URL: https://doi.org/10.11646/zootaxa.5673.3.2

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Additional details

Identifiers

LSID
urn:lsid:zoobank.org:pub:10ACAF18-EDB5-4B60-8299-0FE1D9131E44
LSID
urn:lsid:plazi.org:pub:FF85B607FFDBFFEAFFF4FFF3E211966E
URL
http://publication.plazi.org/id/FF85B607FFDBFFEAFFF4FFF3E211966E

Cites
Publication: 10.1016/j.ympev.2016.07.029 (DOI)
Publication: 10.1111/ecog.03199 (DOI)
Publication: 10.1046/j.1365-294x.2003.01751.x (DOI)
Publication: 10.1016/j.ympev.2006.09.009 (DOI)
Publication: 10.1111/j.0014-3820.2000.tb01253.x (DOI)
Publication: 10.1016/j.ympev.2017.03.017 (DOI)
Publication: 10.11646/zootaxa.3881.6.3 (DOI)
Publication: 10.11646/zootaxa.5154.2.4 (DOI)
Publication: 10.1163/156853809787392711 (DOI)
Publication: 10.3897/herpetozoa.37.e137394 (DOI)
Publication: 10.3897/zookeys.1153.97139 (DOI)
Publication: 10.1093/bioinformatics/17.8.754 (DOI)
Publication: 10.11646/zootaxa.5474.4.3 (DOI)
Publication: 10.1093/molbev/msac052 (DOI)
Publication: 10.1093/bib/bbx108 (DOI)
Publication: 10.11646/zootaxa.5406.1.4 (DOI)
Publication: 10.11646/zootaxa.5380.4.1 (DOI)
Publication: 10.1093/molbev/mss020 (DOI)
Publication: 10.1016/j.ympev.2005.07.016 (DOI)
Publication: 10.11646/zootaxa.5405.2.6 (DOI)
Publication: 10.1016/j.ympev.2020.106807 (DOI)
Publication: 10.1016/j.ympev.2010.04.032 (DOI)
Publication: 10.11646/zootaxa.5570.3.3 (DOI)
Publication: 10.24272/j.issn.2095-8137.2021.085 (DOI)
Publication: 10.5962/bhl.title.12104 (DOI)
Publication: 10.1080/00222933.2019.1656784 (DOI)
Publication: 10.30906/1026-2296-2019-26-6-335-348 (DOI)
Publication: 10.7717/peerj.12713 (DOI)
Publication: 10.30906/1026-2296-2023-30-5-255-476 (DOI)
Publication: 10.30906/1026-2296-2021-28-3A-1-110 (DOI)
Publication: 10.47605/tapro.v9i2.228 (DOI)
Publication: 10.24272/j.issn.2095-8137.2021.228 (DOI)
Publication: 10.1093/sysbio/syy032 (DOI)
Publication: 10.1111/zsc.12007 (DOI)
Publication: 10.1016/j.ympev.2024.108062 (DOI)
Publication: 10.3897/zookeys.1168.95833 (DOI)
Publication: 10.1093/molbev/mst197 (DOI)
Publication: 10.1046/j.1365-2699.2000.00489.x (DOI)
Publication: 10.3390/d16090576 (DOI)
Publication: 10.1201/b16901 (DOI)
Publication: 10.1111/1755-0998.13784 (DOI)
Publication: 10.1007/s10584-008-9457-2 (DOI)
Publication: 10.1371/journal.pone.0151941 (DOI)
Publication: 10.11646/zootaxa.1493.1.1 (DOI)
Has part
Taxonomic treatment: 10.5281/zenodo.17020154 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03BCCE7FFFDDFFFAFF63FE60E09094EB (URL)
Taxonomic treatment: 10.5281/zenodo.17020060 (DOI)
Taxonomic treatment: http://treatment.plazi.org/id/03BCCE7FFFCBFFFAFF63FD25E78D93A4 (URL)
Figure: 10.5281/zenodo.16982239 (DOI)
Figure: 10.5281/zenodo.16982245 (DOI)
Figure: 10.5281/zenodo.16982247 (DOI)
Figure: 10.5281/zenodo.16982253 (DOI)
Dataset: http://table.plazi.org/id/DF6A2FE1FFD2FFE3FF6CFCD3E64D9532 (URL)
Figure: 10.5281/zenodo.16982255 (DOI)
Figure: 10.5281/zenodo.16984280 (DOI)
Dataset: http://table.plazi.org/id/DF6A2FE1FFD7FFE6FF6CFE2FE7639452 (URL)
Figure: 10.5281/zenodo.16982263 (DOI)
Dataset: http://table.plazi.org/id/DF6A2FE1FFCDFFFCFF57FF50E3A196D5 (URL)
Dataset: http://table.plazi.org/id/DF6A2FE1FFCCFFFDFF57FF50E3A196D5 (URL)
Dataset: http://table.plazi.org/id/DF6A2FE1FFC1FFF0FF57FF50E3A196D5 (URL)
Is source of
Dataset: https://www.gbif.org/dataset/1c38160b-7c00-4feb-aa0d-4300c7a31345 (URL)

References

  • Alencar, L.R., Quental, T.B., Grazziotin, F.G., Alfaro, M.L., Martins, M., Venzon, M. & Zaher, H. (2016) Diversification in vipers: phylogenetic relationships, time of divergence and shifts in speciation rates. Molecular Phylogenetics and Evolution, 105, 50-62. https://doi.org/10.1016/j.ympev.2016.07.029
  • Ali, J.R. (2018) New explanation for elements of Hainan Island's biological assemblage may stretch things a little too far. Ecography, 41 (3), 457-460. https://doi.org/10.1111/ecog.03199
  • Balakrishnan, C.N., Monfort, S.L., Gaur, A., Singh, L. & Sorenson, M.D. (2003) Phylogeography and conservation genetics of Eld's deer (Cervus eldi). Molecular Ecology, 12, 1-10. https://doi.org/10.1046/j.1365-294x.2003.01751.x
  • Blyth, E. (1854) Notices and descriptions of various reptiles, new or little known, Part 2. Journal of the Asiatic Society of Bengal, 23 (3), 287-302.
  • Boulenger, G.A. (1894) Catalogue of the snakes in the British Museum (Natural History). Volume II., containing the conclusion of the Colubridae Aglyphae. British Museum (Natural History), London, 382 pp.
  • Burbrink, F.T. & Lawson, R. (2007) How and when did Old World rat snakes disperse into the New World? Molecular Phylogenetics and Evolution, 43 (1), 173-189. https://doi.org/10.1016/j.ympev.2006.09.009
  • Burbrink, F.T., Lawson, R. & Slowinski, J.B. (2000) Mitochondrial DNA phylogeography of the polytypic North American rat snake (Elaphe obsoleta): a critique of the subspecies concept. Evolution, 54, 2107-2118. https://doi.org/10.1111/j.0014-3820.2000.tb01253.x
  • Chan-ard, T., Nabhitabhata, J. & Parr, J.W. (2015) A field guide to the reptiles of Thailand. Oxford University Press, New York, New York, 352 pp.
  • Chen, X., Lemmon, A.R., Lemmon, E.M., Pyron, A.R. & Burbrink, F.T. (2017) Using phylogenomics to understand the link between biogeographic origins and regional diversification in ratsnakes. Molecular Phylogenetics and Evolution, 111, 206-218. https://doi.org/10.1016/j.ympev.2017.03.017
  • Chen, X., McKelvy, A.D., Grismer, L.L., Matsui, M., Nishikawa, K. & Burbrink, F.T. (2014) The phylogenetic position and taxonomic status of the Rainbow Tree Snake Gonyophis margaritatus (Peters, 1871) (Squamata: Colubridae). Zootaxa, 3881 (6), 532-548. https://doi.org/10.11646/zootaxa.3881.6.3
  • David, P., Campbell, P.D., Deuti, K., Hauser, S., Luu, V.Q., Nguyen, T.Q., Orlov, N., Pauwels, O.S.G., Scheinberg, L.R., Sethy, P.G.S., Smits, T., Teynie, A. & Vogel, G. (2022) On the distribution of Gonyosoma prasinum (Blyth, 1854) and Gonyosoma coeruleum Liu, Hou, Ye Htet Lwin, Wang & Rao, 2021, with a note on the status of Gonyosoma gramineum Gunther, 1864 (Squamata: Serpentes: Colubridae). Zootaxa, 5154 (2), 175-197. https://doi.org/10.11646/zootaxa.5154.2.4
  • David, P., Teynie, A. & Vogel, G. (2023) The Snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp.
  • Dowling, H. (1951) A proposed standard system of counting ventrals in snakes. British Journal of Herpetology, 11, 97-99.
  • Gong, S.P., Shi, H.T., Mo, Y.M., Auer, M., Vargas-Ramirez, M., Hundsdorfer, A.K. & Fritz, U. (2009) Phylogeography of the endangered black-breasted leaf turtle (Geoemyda spengleri) and conservation implications for other chelonians. Amphibia-Reptilia, 20, 57-62. https://doi.org/10.1163/156853809787392711
  • Gorin, V.A., Orlov, N.L., Bragin, A.M., Pawangkhanant, P., Milto, K.D., Le, D.X., Nguyen, T.V., Suwannapoom, C. & Poyarkov, N.A. (2024) Phylogeographic pattern and taxonomic revision of the Kaloula baleata species complex (Amphibia: Anura: Microhylidae) with description of two new species from Indochina. Herpetozoa, 37, 391-420. https://doi.org/10.3897/herpetozoa.37.e137394
  • Hall, T.A. (1999) BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symposium Series, 41, 95-98.
  • Hecht, V.L., Pham, C.T., Nguyen, T.T., Nguyen, T.Q., Bonkowski, M. & Ziegler, T. (2013) First report on the herpetofauna of Tay Yen Tu Nature Reserve, northeastern Vietnam. Biodiversity Journal, 4 (4), 507-552.
  • Hu, J., Lan, C. & Li, C. (2023) Microdous amblyrhynchos sp. nov., a new member of the small-toothed sleepers (Teleostei, Gobiiformes, Odontobutidae) from Guangxi, southern China. ZooKeys, 1153, 1-13. https://doi.org/10.3897/zookeys.1153.97139
  • Huelsenbeck, J.P. & Ronquist, F. (2001) MRBAYES: Bayesian inference of phylogenetic trees. Bioinformatics, 17, 754-755. https://doi.org/10.1093/bioinformatics/17.8.754
  • Idiiatullina, S.S., Nguyen, T.V., Bragin, A.M., Pawangkhanant, P., Le, D.X., Vogel, G., David, P. & Poyarkov, N.A. (2024) A new species of green pitviper of the Trimeresurus macrops complex (Reptilia: Serpentes: Viperidae) from the South Central Coast region of Vietnam. Zootaxa, 5474 (4), 375-411. https://doi.org/10.11646/zootaxa.5474.4.3
  • IUCN Standards and Petitions Subcommittee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee, IUCN, Gland, 113 pp.
  • Kambayashi, C., Kakehashi, R., Sato, Y., Mizuno, H., Tanabe, H., Rakotoarison, A., Kunzel, S., Furuno, N., Ohshima, K., Kumazawa, Y., Nagy, Z.T., Mori, A., Allison, A., Donnellan, S.C., Ota, H., Hoso, M., Yanagida, T., Sato, H., Vences, M. & Kurabayashi, A. (2022) Geography-dependent horizontal gene transfer from vertebrate predators to their prey. Molecular Biology and Evolution, 39 (4), msac052. https://doi.org/10.1093/molbev/msac052
  • Katoh, K., Rozewicki, J. & Yamada, K.D. (2019) MAFFT online service: multiple sequence alignment, interactive sequence choice and visualization. Briefings in Bioinformatics, 20, 1160-1166. https://doi.org/10.1093/bib/bbx108
  • Kliukin, N.S., Bragin, A.M., Nguyen, T.V., Le, S.X., Tran, T.T.V., Gorin, V.A. & Poyarkov, N.A. (2024) Another new species of Dibamus Dumeril & Bibron, 1839 (Squamata: Dibamidae) from Nui Chua National Park, Ninh Thuan Province, Vietnam. Zootaxa, 5406 (1), 87-104. https://doi.org/10.11646/zootaxa.5406.1.4
  • Kliukin, N.S., Nguyen, T.V., Le, S.X., Bragin, A.M., Tran, T.T.V., Gorin, V.A. & Poyarkov, N.A. (2023) A new species of the genus Dibamus Dumeril & Bibron, 1839 (Squamata: Dibamidae) from the driest and hottest place of Vietnam. Zootaxa, 5380 (4), 301-320. https://doi.org/10.11646/zootaxa.5380.4.1
  • Lalrinsanga, H.T., Lalremsanga, L., Ht. Decemson, V., Mathipi, L., Tanpuii, L., Muansanga, L. & Biakzuala, L. (2022) Contributions to the morphology and molecular phylogenetics of Gonyosoma prasinum (Blyth, 1854) (Reptilia: Squamata: Colubridae) from Mizoram, India. Hamadryad, 39, 96-103.
  • Lanfear, R., Calcott, B., Ho, S.Y.W. & Guindon, S. (2012) PartitionFinder: combined selection of partitioning schemes and substitution models for phylogenetic analyses. Molecular Biology and Evolution, 29, 1695-1701. https://doi.org/10.1093/molbev/mss020
  • Lawson, R., Slowinski, J.B., Crother, B.I. & Burbrink, F.T. (2005) Phylogeny of the Colubroidea (Serpentes): new evidence from mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution, 37 (2), 581-601. https://doi.org/10.1016/j.ympev.2005.07.016
  • Li, H.Z., Liu, Z. & Li, J.L. (2024) A new species of the genus Celaenorrhinus Hubner, [1819] from Hainan, China (Lepidoptera, Hesperiidae). Zootaxa, 5405 (2), 265-272. https://doi.org/10.11646/zootaxa.5405.2.6
  • Li, J.N., Liang, D., Wang, Y.Y., Guo, P., Huang, S. & Zhang, P. (2020) A large-scale systematic framework of Chinese snakes based on a unified multilocus marker system. Molecular Phylogenetics and Evolution, 148, 106807. https://doi.org/10.1016/j.ympev.2020.106807
  • Liang, G. (2018) A study of the genesis of Hainan Island. Geology in China, 45, 693-705. [in Chinese]
  • Lin, L.H., Ji, X., Diong, C.H., Du, Y. & Lin, C.X. (2010) Phylogeography and population structure of the Reevese's Butterfly Lizard (Leiolepis reevesii) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 56, 601- 607. https://doi.org/10.1016/j.ympev.2010.04.032
  • Luu, V.Q., Dinh, T.S., Lo, O.V., Nguyen, T.Q. & Ziegler, T. (2020) New records and an updated list of reptiles from Ba Vi National Park, Vietnam. Bonn Zoological Bulletin, 69 (1), 1-9.
  • Messenger, K.R. (2021) The Asian Ratsnakes and Kin of Greater China. Privately printed by the author, Nanjing Forestry University, Nanjing, xv + 174 pp.
  • Nguyen, S.V., Ho, C.T. & Nguyen, T.Q. (2009) Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, 768 pp.
  • Nguyen, T.V. & Vogel, G. (2024) A new species of Lycodon Fitzinger, 1826 from the northern part of the Truong Son Mountains, Central Vietnam (Reptilia: Squamata: Colubridae). Zootaxa, 5570 (3), 484-510. https://doi.org/10.11646/zootaxa.5570.3.3
  • Nguyen, T.V., Brakels, P., Maury, N., Sudavanh, S., Pawangkhanant, P., Idiiatullina, S., Lorphengsy, S., Inkhavilay, K., Suwannapoom, C. & Poyarkov, N.A. (2020) New herpetofaunal observations from Laos based on photo records. Amphibian & Reptile Conservation, 14 (2), e248, 218-249.
  • Peng, L.F., Zhang, Y., Huang, S., Burbrink, F.T., Chen, J.M., Hou, M., Zhu, Y.W., Yang, H. & Wang, J.C. (2021) A new snake species of the genus Gonyosoma Wagler, 1828 (Serpentes: Colubridae) from Hainan Island, China. Zoological Research, 42 (4), 487-491. https://doi.org/10.24272/j.issn.2095-8137.2021.085
  • Pope, C.H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of Central Asia, Vol. X. American Museum of Natural History, New York, New York, 604 pp. https://doi.org/10.5962/bhl.title.12104
  • Poyarkov, N.A., Nguyen, T.V. & Vogel, G. (2019) A new species of the genus Liopeltis Fitzinger, 1843 from Vietnam (Squamata: Colubridae). Journal of Natural History, 53 (27-28), 1647-1672. https://doi.org/10.1080/00222933.2019.1656784
  • Poyarkov, N.A., Nguyen, T.V., Orlov, N.L. & Vogel, G. (2019) A new species of the genus Calamaria Boie, 1827 from the highlands of the Langbian Plateau, southern Vietnam (Squamata: Colubridae). Russian Journal of Herpetology, 26 (6), 335-348. https://doi.org/10.30906/1026-2296-2019-26-6-335-348
  • Poyarkov, N.A., Nguyen, T.V., Pawangkhanant, P., Yushchenko, P.V., Brakels, P., Nguyen, L.H., Nguyen, H.N., Suwannapoom, C., Orlov, N.L. & Gernot, V. (2022) An integrative taxonomic revision of Pareineae slug-eating snakes (Squamata: Pareidae) reveals an unprecedented diversity in Indochina. PeerJ, 10, e12713. https://doi.org/10.7717/peerj.12713
  • Poyarkov, N.A., Nguyen, T.V., Popov, E.S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C. & Orlov, N.L. (2023) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of reptiles in Indochina. Russian Journal of Herpetology, 30 (5), 255-476. https://doi.org/10.30906/1026-2296-2023-30-5-255-476
  • Poyarkov, N.A., Nguyen, T.V., Popov, E.S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C. & Orlov, N.L. (2021) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of amphibians in Indochina. Russian Journal of Herpetology, 28 (3A), 1-110. https://doi.org/10.30906/1026-2296-2021-28-3A-1-110
  • Poyarkov, N.A., Nguyen, T.V., Trofimets, A.V. & Gorin, V.A. (2020) A new cryptic species of the genus Microhyla (Amphibia: Microhylidae) from Langbian Plateau, Vietnam. Taprobanica, 9 (2), 136-163. https://doi.org/10.47605/tapro.v9i2.228
  • Poyarkov, N.A., Nguyen, T.V., Yang, J.H. & Gorin, V. (2021) A new species of Micryletta (Amphibia: Anura: Microhylidae) from Langbian Plateau in southern Vietnam. Zoological Research, 42 (6), 726-733. https://doi.org/10.24272/j.issn.2095-8137.2021.228
  • Pham, A.V., Nguyen, S.L.H. & Nguyen, T.Q. (2014) New records of snakes (Squamata: Serpentes) from Son La Province, Vietnam. Herpetology Notes, 7, 771-777.
  • Rambaut, A., Drummond, A.J., Xie, D., Baele, G. & Suchard, M.A. (2018) Posterior summarization in Bayesian phylogenetics using Tracer 1.7. Systematic Biology, 67, 901-904. https://doi.org/10.1093/sysbio/syy032
  • Sambrook, J., Fritsch, E.F. & Maniatis, T. (1989) Molecular Cloning: A Laboratory Manual. 3 Vols. Cold Spring Harbor Laboratory Press, Cold Spring Harbor, 1625S.
  • Schulz, K.D. (1996) A Monograph of the Colubrid Snakes of the Genus Elaphe Fitzinger. Koeltz Scientific Books, Havlickuv Brod, iii + 439 pp.
  • Schulz, K.D. (2013) An annotated and illustrated checklist of Old World ratsnakes. In: Schulz, K.D. (Ed.), Old World Ratsnakes - A Collection of Papers. Bushmaster Publications, Berg, pp. 17-268.
  • Shi, H.T., Zhao, E.M., Wang, L.J., Bi, H., Lyu, S.Q., Liu, H.N., Wang, J.C., Zhao, H. & Hong, M.L. (2011) Amphibian and Reptile Fauna of Hainan. Science Press, Beijing, 285 pp.
  • Shi, Y.F., Cui, Z.J. & Su, Z. (2006) The Quaternary Glaciations and Environmental Variations in China. Hebei Science and Technology Press, Hebei, 618 pp.
  • Siler, C.D., Oliveros, C.H., Santanen, A. & Brown, R.M. (2013) Multilocus phylogeny reveals unexpected diversification patterns in Asian wolf snakes (genus Lycodon). Zoologica Scripta, 42 (3), 262-277. https://doi.org/10.1111/zsc.12007
  • Smith, M.A. (1943) The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor & Francis, London, 583 pp.
  • Stuart, B.L. & Heatwole, H. (2008) Country records of snakes from Laos. Hamadryad, 33, 97-106.
  • Stuart, B.L. (1999) Amphibians and reptiles. In: Duckworth, J.W., Salter, R.E. & Khounboline, K. (Eds.), Wildlife in Lao PDR: 1999 Status Report. IUCN, Wildlife Conservation Society, Centre for Protected Area and Watershed, Vientiane, pp. 43-67.
  • Sun, X.Y., Pan, Q.Q., Hubley, B., Ye, Z., Zhang, P. & Xie, Q. (2024) Geomorphic impacts within Red River Fault and island shifting as witnessed by the phylogeography of the largest water strider. Molecular Phylogenetics and Evolution, 195, 108062. https://doi.org/10.1016/j.ympev.2024.108062
  • Syafiq, M.F., Badli-Sham, B.H., Grismer, L.L. & Ahmad, A.B. (2023) Uneven species occurrence and richness of lowland snakes (Serpentes, Squamata) in Terengganu, Peninsular Malaysia, with new locality records. ZooKeys, 1168, 11-39. https://doi.org/10.3897/zookeys.1168.95833
  • Tamura, K., Stecher, G., Peterson, D., Filipski, A. & Kumar, S. (2013) MEGA6: Molecular Evolutionary Genetics Analysis version 6.0. Molecular Biology and Evolution, 30, 2725-2729. https://doi.org/10.1093/molbev/mst197
  • Tweedie, M.W.F. (1983) The Snakes of Malaya. 3rd Edition. Singapore National Printers, Singapore, 167 pp., pls. 1-12.
  • Uetz, P., Freed, P., Aguilar, R. & Hosek, J. (2025) The Reptile Database. Available from: http://www.reptile-database.org/ (accessed 1 March 2025)
  • Voris, H.K. (2000) Maps of Pleistocene sea levels in Southeast Asia: shorelines, river systems and time durations. Journal of Biogeography, 27, 1153-1167. https://doi.org/10.1046/j.1365-2699.2000.00489.x
  • Wallach, V., Midtgaard, R. & Hsiao, E. (2024) Revalidation of the arboreal Asian snake genera Gonyophis Boulenger, 1891, Rhynchophis Mocquard, 1897, and Rhadinophis Vogt, 1922, with description of a new genus and tribe (Squamata: Serpentes: Colubridae). Diversity, 16, 576. https://doi.org/10.3390/d16090576
  • Wallach, V., Williams, K.L. & Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, Florida, xxvii + 1209 pp. https://doi.org/10.1201/b16901
  • Wangyal, J.T., Sherub, D.S.B., Tshewang, S., Wangdi, D., Rinchen, K., Puntsho, S., Tashi, C., Koirala, B.K., Bhandari, G.S., Phuntsho, Y., Koirala, T.P., Ghalley, B.B., Chaida, L., Tenzin, J., Powrel, R.B., Tshewang, R., Raika, O.N., Jamtsho, S., Kinley, G., Gyeltshen, T., Tashi, S., Nidup, D., Phuentsho, N.W., Norbu, L., Wangdi, K., Wangchuk, T., Tobgay, P., Dorji, T. & Das, I. (2020) New herpetofaunal records from the Kingdom of Bhutan obtained through citizen science. Herpetological Review, 51 (4), 790-798.
  • Wu, G.J., Wu, T.H., Zhu, J. & Xiao, L. (2002) A new record of snake in Sichuan -Elaphe prasina. Sichuan Journal of Zoology, 21 (2), 66. [in Chinese]
  • Wu, Y.H., Hou, S.B., Yuan, Z.Y., Jiang, K., Huang, R.Y., Wang, K., Liu, Q., Yu, Z.B., Zhao, H.P., Zhang, B.L., Chen, J.M., Wang, L.J., Stuart, B.L., Chambers, E.A., Wang, Y.F., Gao, W., Zou, D.H., Yan, F., Zhao, G.G., Fu, Z.X., Wang, S.N., Jiang, M., Zhang, L., Ren, J.L., Wu, Y.Y., Zhang, L.Y., Yang, D.C., Jin, J.Q., Yin, T.T., Li, J.T., Zhao, W.G., Murphy, R.W., Huang, S., Guo, P., Zhang, Y.P. & Che, J. (2023) DNA barcoding of Chinese snakes reveals hidden diversity and conservation needs. Molecular ecology resources, 23, 1124-1141. https://doi.org/10.1111/1755-0998.13784
  • Yang, D. & Rao, D.Q. (2008) Amphibia and Reptilia of Yunnan. Yunnan Publishing Group Corporation, Yunnan Science and Technology Press, Kunming, 8 + 411 pp. [in Chinese]
  • Yao, Y.F., Bera, S., Ferguson, D.K., Mosbrugger, V., Paudayal, K.N., Jin, J.H. & Li, C.S. (2009) Reconstruction of paleovegetation and paleoclimate in the Early and Middle Eocene, Hainan Island, China. Climatic Change, 92, 169-189. https://doi.org/10.1007/s10584-008-9457-2
  • Zhao, E.M., Huang, M.H. & Zong, Y. (1998) Fauna Sinica: Reptilia, Volume 3. Squamata, Serpentes. Science Press, Beijing, 570 pp.
  • Zhao, H.T., Wang, L.R. & Yuan, J.Y. (2007) Origin and time of Qiongzhou Strait. Marine Geology and Quaternary Geology, 27, 33-40.
  • Zhu, H. (2016) Biogeographical evidences help revealing the origin of Hainan Island. PLoS One, 11 (4), e0151941. https://doi.org/10.1371/journal.pone.0151941
  • Ziegler, T., Hendrix, R., Vu, T.N., Vogt, M., Forster, B. & Dang, K.N. (2007) The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa, 1493 (1), 1-40. https://doi.org/10.11646/zootaxa.1493.1.1