Published September 28, 2023 | Version v1
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New sauropod dinosaur from the Lower Cretaceous of Morella (Spain) provides new insights on the evolutionary history of Iberian somphospondylan titanosauriforms

  • 1. Instituto Dom Luiz, Universidade de Lisboa, Edifício C6, Campo Grande, 1749-016 Lisboa, Portugal
  • 2. Grupo de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional de Educación a Distancia (UNED), Avda. Esparta s/n, 28232 Las Rozas de Madrid, Spain
  • 3. Institut Català de Palaeontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Carrer Escola Industrial 23, 08201 Sabadell, Spain
  • 4. Grup Guix, C/ Santa Lucia, 75, E-12540 Vila-real, Spain & Museo de Ciencias Naturales, C/ General Elio s/n, E-46010 València, Spain
  • 5. Grup Guix, C/ Santa Lucia, 75, E-12540 Vila-real, Spain & Àrea de Cristal•lografia i Mineralogia, Departament de Biologia, Bioquímica i Ciències Naturals, Universitat Jaume I, Av/de Vicent Sos Baynat, s/n, E-12071 Castelló de la Plana, Spain
  • 6. Grupo de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional de Educación a Distancia (UNED), Avda. Esparta s/n, 28232 Las Rozas de Madrid, Spain & Unidad de Palaeontología, Universidad Autónoma de Madrid, C/ Darwin, 2, 28049 Madrid, Spain & Real Academia de Ciencias Exactas, Físicas y Naturales, C/ de Valverde, 24, 28004 Madrid, Spain

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Mocho, Pedro, Escaso, Fernando, Gasulla, José M., Galobart, Àngel, Poza, Begoña, Santos-Cubedo, Andrés, Sanz, José L., Ortega, Francisco (2024): New sauropod dinosaur from the Lower Cretaceous of Morella (Spain) provides new insights on the evolutionary history of Iberian somphospondylan titanosauriforms. Zoological Journal of the Linnean Society 201 (201): 214-268, DOI: 10.1093/zoolinnean/zlad124, URL: https://www.mendeley.com/catalogue/76af828d-3d52-3fc1-8f55-e11c3bdf1b2c/

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References

  • Allain R, Taquet P, Battail B et al. Un nouveau genre de dinosaure sauropode de la formation de Gres superieurs (Aptien-Albien) du Laos. Comptes Rendus de l'Academie des Sciences de Paris, Sciences de la Terres et des Planetes 1999;329:609-16.
  • Allain R, Vullo R, Le Loeuff J et al. European ornithomimosaurs (Dinosauria, Theropoda): an undetected record. Geologica Acta 2014;12:127-35.
  • Allain R, Vullo R, Rozada L et al. Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstatte (southwestern France): implications for continental faunal turnover at the J/K boundary. Geodiversitas 2022;44:683-752.
  • Bellardini F, Coria RA, Pino DA et al. Osteology and phylogenetic relationships of Ligabuesaurus leanzai (Dinosauria: Sauropoda) from the Early Cretaceous of the Neuquen Basin, Patagonia, Argentina. Zoological Journal of the Linnean Society 2022;196:1333-93.
  • Bonaparte JF, Heinrich W-D, Wild R. Review of Janenschia Wild, with the description of a new sauropod from the Tendaguru beds of Tanzania and a discussion on the systematic value of procoelous caudal vertebrae in the Sauropoda. Palaeontographica, Abteilung A 2000;256:25-76
  • Bonaparte JF, Gonzalez Riga BJ, Apesteguia S. Ligabuesaurus leanzai gen. et sp. nov. (Dinosauria, Sauropoda), a new titanosaur from the Lohan Cura Formation (Aptian, Lower Cretaceous) of Neuquen, Patagonia, Argentina. Cretaceous Research 2006;27:364-76.
  • Borsuk-Bialynicka M. A new camarasaurid sauropod Opisthocoelicaudia skarzynskii gen. n., sp. n. from the Upper Cretaceous of Mongolia. Palaeontologica Polonica 1977;37:5-63.
  • Bover-Arnal T, Moreno-Bedmar JA, Frijia G et al. Chronostratigraphy of the Barremian-Early Albian of the Maestrat Basin (E Iberian Peninsula): integrating strontium-isotope stratigraphy and ammonoid biostratigraphy. Newsletters on Stratigraphy 2016;49:41-68.
  • Brikiatis L. Late Mesozoic North Atlantic land bridges. Earth-Science Reviews 2016;159:47-57.
  • Britt BB, Scheetz RD, Whiting MF et al. Moabosaurus utahensis, n. gen., n. sp., a new sauropod from the Early Cretaceous (Aptian) of North America. Contributions from the Museum of Paleontology University of Michigan 2017;32:189-243.
  • Buffetaut E, Azar D, Nel A et al. First nonavian dinosaur from Lebanon: a brachiosaurid sauropod from the Lower Cretaceous of the Jezzine District. Naturwissenschaften 2006;93:440-3.
  • Buscalioni AD, Poyato-Ariza FJ. Las Hoyas: a unique Cretaceous ecosystem. New Mexico Museum of Natural History and Science Bulletin 2016;71:49-61.
  • Canudo JI, Ruiz-Omenaca JI, Barco JL et al. ?Sauropodos asiaticos en el Barremiense inferior (Cretacico Inferior) de Espana? Ameghiniana 2002;39:443-52.
  • Canudo JI, Ruiz-Omenaca JI, Sender LM. Primera evidencia de un dinosaurio sauropodo en la Formacion Escucha (Utrillas, Teruel), Albiense medio (Cretacico inferior). Geo-Temas 2004a;6:27-30.
  • Canudo JI, Ruiz-Omenaca JI, Del Ramo A et al. Primera evidencia de restos de dinosaurio en Murcia (Cretacico Inferior, Albiense). Geogaceta 2004b;35:119-22.
  • Canudo JI, Royo-Torres R, Cuenca-Bescos G. A new sauropod: Tastavinsaurus sanzi gen. et sp. nov. from the Early Cretaceous (Aptian) of Spain. Journal of Vertebrate Paleontology 2008;28:712-31.
  • Canudo JI, Barco JL, Pereda-Suberbiola X et al. What Iberian dinosaurs reveal about the bridge said to exist between Gondwana and Laurasia in the Early Cretaceous. Bulletin de la Societe Geologique de France 2009;180:5-11.
  • Canudo JI, Carballido JL, Garrido A et al. A new rebbachisaurid sauropod from the Aptian-Albian, Lower Creta ceous Rayoso Formation, Neuquen, Argentina. Acta Palaeontologica Polonica 2018;63:679-91.
  • Carballido JL, Sander PM. Postcranial axial skeleton of Europasaurus holgeri (Dinosauria, Sauropoda) from Upper Jurassic of Germany: implications for sauropod ontogeny and phylogenetic relationships of basal Macronaria. Journal of Systematic Paleontology 2014;12:335-87.
  • Carballido JL, Pol D, Cerda I et al. The osteology of Chubutisaurus insignis del Corro, 1975 (Dinosauria: Neosauropoda) from the 'middle' Cretaceous of central Patagonia, Argentina. Journal of Vertebrate Paleontology 2011a;31:93-110.
  • Carballido JL, Rauhut OWM, Pol D et al. Osteology and phylogeny relationships of Tehuelchesaurus benitezii (Dinosauria, Sauropoda) from the Upper Jurassic of Patagonia. Zoological Journal of the Linnean Society 2011b;163:605-62.
  • Carballido JL, Salgado L, Pol D et al. A new basal rebbachisaurid (Sauropoda, Diplodocoidea) from the Early Cretaceous of the Neuquen Basin; evolution and biogeography of the group. Historical Biology 2012;24:631-54.
  • Carballido JL, Pol D, Parra Ruge ML et al. A new Early Cretaceous brachiosaurid (Dinosauria, Neosauropoda) from north-western Gondwana (Villa de Leiva, Colombia). Journal of Vertebrate Paleontology 2015;35:e980505.
  • Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings of the Royal Society B: Biological Sciences 2017;284:20171219.
  • Carballido JL, Scheil M, Knotschke N et al. The appendicular skeleton of the dwarf macronarian sauropod Europasaurus holgeri from the Late Jurassic of Germany and a re-evaluation of its systematic affinities. Journal of Systematic Palaeontology 2020;18:739-81.
  • Carrano MT. The evolution of sauropod locomotion. In: Curry Rogers D, Wilson JA (eds), The Sauropods, Evolution and Paleobiology. Berkeley: University of California Press, 2005, 229-51.
  • Casanovas ML, Ezquerra R, Fernandez A et al. Dos nuevos yacimientos de icnitas de dinosaurios en la Rioja y en la provincia de Soria (Espana). Coloquios de Paleontologia 1995;47:9-23.
  • Casanovas ML, Fernandez A, Perez-Lorente F et al. Sauropod trackways from site El Sobaquillo (Munilla, La Rioja, Spain) indicate amble walking. Ichnos 1997;5:101-7.
  • Chure D, Britt BB, Whitlock JA et al. First complete sauropod dinosaur skull from the Cretaceous of the Americas and the evolution of sauropod dentition. Naturwissenschaften 2010;97:379-91.
  • Congreve CR, Lamsdell JC. Implied weighting and its utility in palaeontological datasets: a study using modelled phylogenetic matrices. Palaeontology 2016;59:447-62.
  • Coria RA, Windholz GJ, Ortega F et al. A new dicraeosaurid sauropod from the Lower Cretaceous (Mulichinco Formation, Valanginian, Neuquen Basin) of Argentina. Cretaceous Research 2019;93:33-48.
  • Cuenca-Bescos G, Canudo JI. A new gobiconodontid mammal from the Early Cretaceous of Spain and its palaeogeographic implications. Acta Palaeontologica Polonica 2003;48:575-82.
  • Curry Rogers K. The postcranial osteology of Rapetosaurus krausei (Sauropoda: Titanosauria) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology 2009;29:1046-86.
  • Dal Sasso C, Pierangelini G, Famiani F et al. First sauropod bones from Italy offer new insights on the radiation of Titanosauria between Africa and Europe. Cretaceous Research 2016;64:88-109.
  • D'Emic MD. The early evolution of titanosauriform sauropod dinosaurs. Zoological Journal of the Linnean Society 2012;166:624-71.
  • D'Emic MD. Revision of the sauropod dinosaurs of the Early Cretaceous Trinity Group, southern USA, with the description of a new genus. Journal of Systematic Palaeontology 2013;11:707-26.
  • D'Emic MD, Foreman BZ. The beginning of the sauropod dinosaur hiatus in North America: insights from the Lower Cretaceous Cloverly Formation of Wyoming. Journal of Vertebrate Paleontology 2012;32:883-902.
  • D'Emic MD, Wilson JA, Williamson TE. A sauropod dinosaur pes from the latest Cretaceous of North America and the validity of Alamosaurus sanjuanensis (Sauropoda, Titanosauria). Journal of Vertebrate Paleontology 2011;31:1072-9.
  • D'Emic MD, Mannion PD, Upchurch P et al. Osteology of Huabeisaurus allocotus (Sauropoda:Titanosauriformes) from the Upper Cretaceous of China. PLoS One 2013;8:e69375.
  • D'Emic MD, Foreman BZ, Jud NA. Anatomy, systematics, paleoenvironment, growth, and age of the sauropod dinosaur Sonorasaurus thompsoni from the Cretaceous of Arizona, USA. Journal of Paleontology 2016;90:102-32.
  • Diez Diaz V, Pereda Suberbiola X, Sanz JL. The axial skeleton of the titanosaur Lirainosaurus astibiae (Dinosauria: Sauropoda) from the Latest Cretaceous of Spain. Cretaceous Research 2013a;43:145-60.
  • Diez Diaz V, Pereda Suberbiola X, Sanz JL. Appendicular skeleton and dermal armour of the Late Cretaceous titanosaur Lirainosaurus astibiae (Dinosauria:Sauropoda) from Spain. Palaeontologia Electronica 2013b;16.2.19A:18.
  • Diez Diaz V, Mocho P, Paramo A et al. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Research 2016;68:49-60.
  • Fuentes Vidarte C, Meijide Calvo M, Meijide Fuentes F et al. Fauna de vertebrados del Cretacico Inferior del Yacimiento de "Zorralbo" en Golmayo (Soria, Espana). Revista Espanola de Paleontologia Numero Extraordinario 2005;X:83-92.
  • Gallina PA, Apesteguia S. Cathartesaura anaerobica gen. et sp. nov., a new rebbachisaurid (Dinosauria, Sauropoda) from the Huincul Formation (Upper Cretaceous), Rio Negro, Argentina. Revista del Museo Argentino de Ciencias Naturales n.s 2005;7:153-66.
  • Gallina PA, Canale JI, Carballido JL. The earliest known titanosaur sauropod dinosaur. Ameghiniana 2021;58:35-51.
  • Gallup MR. Functional morphology of the hindfoot of the Texas sauropod Pleurocoelus sp. indet. In: Farlow JO (ed.), Paleobiology of the Dinosaurs. Boulder: Geological Society of America Special Paper 238, 1989, 71-4.
  • Garcia R, Moreno-Bedmar JA, Bover-Arnal T et al. Lower Cretaceous (Hauterivian-Albian) ammonite biostratigraphy in the Maestrat Basin (E Spain). Journal of Iberian Geology 2014;40:285-312.
  • Gasulla JM, Sanz JL, Escaso F et al. Elementos de la cintura pelvica de dinosaurios sauropodos Titanosauriformes del Cretacico Inferior (Aptiense inferior) de Morella (Castellon). In: Ruiz-Omenaca JI, Pinuela L, Garcia Ramos JC (eds), Libro de resumenes de las 24a Jornadas de la Sociedad Espanola de Paleontologia, Colunga. 2008, 129-30.
  • Gasulla JM, Ortega F, Escaso F et al. Los yacimientos de vertebrados de la Formacion Arcillas de Morella (Aptiense inferior).In: Perez-Garcia A, Gasco F, Gasulla JM, Escaso F (eds), Viajando a Mundos Preteritos. Morella: Ajuntament de Morella, 2011, 157-71.
  • Gasulla JM, Ortega F, Sanz JL et al. Els vertebrats de la Formacio Argiles de Morella (Aptia inferior, Cretaci Inferior). Nemus 2012;2:15-27.
  • Gasulla JM, Escaso F, Narvaez I et al. A new sail-backed styracosternan (Dinosauria: Ornithopoda) from the Early Cretaceous of Morella, Spain. PLoS One 2015;10:e0144167.
  • Gasulla JM, Escaso F, Malafaia E et al. Nuevos yacimientos de vertebrados de la Formacion Arcillas de Morella (Cretacico Inferior, Barremiense) en la Concesion Minera Vega del Moll (Morella,Castellon).In:Garcia-Forner A, Conejero-Ortega N, Diaz-Acha Y, Baratas Diaz A (eds), La huella humana en la naturaleza, Libro de Resumenes, XXIV Bienal de la Real Sociedad Espanola de Historia Natural, Valencia, Septiembre 2021. Madrid: Real Sociedade Espanola de Historia Natural, 2021, 218-9.
  • Gilmore CW. A nearly complete articulated skeleton of Camarasaurus, a saurischian dinosaur from the Dinosaur National Monument, Utah. Memoirs of the Carnegie Museum 1925;10:347-84.
  • Gilmore CW. Osteology of Apatosaurus with special reference to specimens in the Carnegie Museum. Memoirs of the Carnegie Museum 1936;11:175-300.
  • Goloboff PA. Extended implied weighting. Cladistics 2014;30:260-72.
  • Goloboff PA, Catalano SA. TNT v.1.5, including a full implementation of phylogenetic morphometrics. Cladistics 2016;32:221-38.
  • Goloboff PA, Farris JS, Nixon KC. TNT, a free program for phylogenetic analysis. Cladistics 2008;24:774-86.
  • Goloboff PA, Mattoni CI, Quinteros AS. Continuous characters analyzed as such. Cladistics 2006;22:589-601.
  • Goloboff PA, Torres A, Arias JS. Weighted parsimony outperforms other methods of phylogenetic inference under models appropriate for morphology. Cladistics 2018;34:407-37.
  • Ullmann PV, Lacovara KJ. Appendicular osteology of Dreadnoughtus schrani, a giant titanosaurian (Sauropoda, Titanosauria) from the Upper Cretaceous of Patagonia, Argentina. Journal of Vertebrate Paleontology 2016;36:e1225303.https://doi.org/10.1080/02724634.2016.1225303
  • Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi. Palaeontologia Electronica 2005;8:1-37.
  • Gonzalez Riga BJ, Calvo JO, Porfiri J. An articulated titanosaur from Patagonia (Argentina): new evidence of neosauropod pedal evolution. Palaeoworld 2008;17:33-40.
  • Gonzalez Riga BJ, Ortiz David LD, Tomaselli MB et al. Sauropod and theropod dinosaur tracks from the Upper Cretaceous of Mendoza (Argentina): trackmakers and anatomical evidences. Journal of South American Earth Sciences 2015;61:134-41.
  • Gonzalez Riga BJ, Lamanna MC, Ortiz David LD et al. A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot. Scientific Reports 2016;6:19165.
  • Gonzalez Riga BJ, Mannion PD, Poropat SF et al. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 2018;184:136-81.
  • Gorscak E, O'Connor PM. Time-calibrated models support congruency between Cretaceous continental rifting and titanosaurian evolutionary history. Biology Letters 2016;12:20151047.
  • Guimera J, Salas R. Inversion terciaria de la falla normal mesozoica que limitaba la subcuenca de Galve (Cuenca del Maestrazgo). Geogaceta 1996;20:1701-3.
  • Harris JD. The appendicular skeleton of Suuwassea emilieae (Sauropoda: Flagellicaudata) from the Upper Jurassic Morrison Formation of Montana (USA). Geobios 2007;40:501-22.
  • Hatcher JB. Diplodocus (Marsh): its osteology, taxonomy, and probable habits, with a restoration of the skeleton. Memoirs of the Carnegie Museum 1901;1:1-63.
  • He X-L, Li K, Cai K-J. The Middle Jurassic Dinosaur Fauna from Dashanpu, Zigong, Sichuan. Vol IV. Sauropod Dinosaurs (2). Omeisaurus tianfuensis. Chengdu, China: Sichuan Publishing House of Science and Technology, 1988 [in Chinese, English summary].
  • Henderson DM. Burly gaits: centers of mass, stability and the trackways of sauropod dinosaurs. Journal of Vertebrate Paleontology 2006;26:907-21.
  • Huene F. von. Los saurisquios y ornitisquios del Cretacico Argentino. Museo de La Plata, Anales 1929;3:1-194.
  • Hutchinson JR. The evolution of femoral osteology and soft tissues on the line to extant birds (Neornithes). Zoological Journal of the Linnean Society 2001;131:169-97.
  • Janensch W.Die wirbelsaule von Brachiosaurus brancai. Palaeontographica (Supplement VII) 1950;3:27-93.
  • Janensch W. Die Gliedmaszen und Gliedmaszengurtel der Sauropoden der Tendaguru-Schichten. Palaeontographica, Supplementbande 1961;7:177-235.
  • Jannel A, Salisbury SW, Panagiotopoulou O. Softening the steps to gigantism in sauropod dinosaurs through the evolution of a pedal pad. Science Advances 2022;8:eabm8280.
  • Ksepka DT, Norell MA. Erketu ellisoni, a long-necked sauropod from Bor Guve (Dornogov Aimag, Mongolia). American Museum Novitates 2006;3508:1-16.
  • Lacovara KJ, Lamanna MC, Ibiricu LM et al. A gigantic, exceptionally complete titanosaurian sauropod dinosaur from southern Patagonia, Argentina. Scientific Reports 2014;4:1-9.
  • Lapparent AF. Les dinosaures jurassiques de Damparis ( Jura). Memoires de la Societe Geologique de France, Nouvelle Serie 1943;47:1-21.
  • Lapparent AF. Nouveux gisements de reptiles Mesozoiques en Espagne. Notas y Comunicaciones del Instituto Geologico y Minero de Espana 1966;84:103-10.
  • Lapparent AF, Zbyszewski G. Les dinosauriens du Portugal. Memorias dos Servicos Geologicos de Portugal 1957;2:1-63.
  • Lapparent AF, Curnelle R, Defaut B et al. Nouveaux gisements de Dinosaures en Espagne centrale. Estudios Geologicos 1969;25:311-5.
  • Le Loeuff J. Osteology of Ampelosaurus atacis (Titanosauria) from Southern France. In: Tidwell V, Carpenter K (eds), Thunder-Lizards: The Sauropodomorph Dinosaurs. Bloomington and Indianopolis: Indiana University Press, 2005, 115-37.
  • Liao C-C, Moore A, Jin C et al. A possible brachiosaurid (Dinosauria, Sauropoda) from the mid-Cretaceous of northeastern China. PeerJ 2021;9:e11957.
  • Malafaia E, Gasulla JM, Escaso F et al. A new spinosaurid theropod (Dinosauria: Megalosauroidea) from the upper Barremian of Vallibona, Spain: Implications for spinosaurid diversity in the Early Cretaceous of the Iberian Peninsula. Cretaceous Research 2019;106:104221.
  • Mannion PD.A turiasaurian sauropod dinosaur from the Early Cretaceous Wealden Supergroup of the United Kingdom. PeerJ 2019;7:e6348.
  • Mannion PD, Calvo JO. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albian- early Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 2011;163:no-no.
  • Mannion PD, Upchurch P, Barnes RN et al. Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal Titanosauriformes.Zoological Journal of the Linnean Society 2013;168:98-206.
  • Mannion PD, Allain R, Moine O. The earliest known titanosauriform sauropod dinosaur and the evolution of Brachiosauridae. PeerJ 2017;5:e3217.
  • Mannion PD, Upchurch P, Jin X et al. New information on the Cretaceous sauropod dinosaurs of Zhejiang Province, China:impact on Laurasian titanosauriform phylogeny and biogeography. Royal Society Open Science 2019a;6:191057.
  • Mannion PD, Upchurch P, Schwarz D et al. Taxonomic affinities of the putative titanosaurs from the Late Jurassic Tendaguru Formation of Tanzania: phylogenetic and biogeographic implications for eusauropod dinosaur evolution. Zoological Journal of the Linnean Society 2019b;185:784-909.
  • Martin V, Suteethorn V, Buffetaut E. Description of the type and referred material of Phuwiangosaurus sirindhornae Martin, Buffetaut and Suteethorn, 1994, a sauropod from the Lower Cretaceous of Thailand. Oryctos 1999;2:39-91.
  • Martin T, Marugan-Lobon J, Vullo R et al. A Cretaceous eutriconodont and integument evolution in early mammals.Nature 2015;526:380-4.
  • Martinez RD, Gimenez O, Rodriguez J et al. An articulated specimen of the basal titanosaurian (Dinosauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal Formation of Chubut province, Argentina. Journal of Vertebrate Paleontology 2004;24:107-20.
  • Martinez RDF, Lamanna MC, Novas FE et al. A basal lithostrotian titanosaur (Dinosauria: Sauropoda) with a complete skull: implications for the evolution and paleobiology of Titanosauria. PLoS One 2016;11:e0151661.
  • McIntosh JS, Miller WE, Stadtman KL et al. The osteology of Camarasaurus lewisi ( Jensen, 1988). Brigham Young University Geology Studies 1996a;41:73-115.
  • McIntosh JS, Miles CA, Cloward KC et al. A new nearly complete skeleton of Camarasaurus. Bulletin of Gunma Museum of Natural History 1996b;1:1-87.
  • McPhee BW, Mannion PD, Klerk WJ et al. High diversity in the sauropod dinosaur fauna of the Lower Cretaceous Kirkwood Formation of South Africa: implications for the Jurassic-Cretaceous transition. Cretaceous Research 2016;59:228-48.
  • MochoP,Royo-TorresR,OrtegaF.Phylogeneticreassessmentof Lourinhasaurus alenquerensis, a basal Macronaria (Sauropoda) from the Upper Jurassic of Portugal.Zoological Journal of the Linnean Society 2014;170:875-916.
  • Mocho P, Escaso F, Gasulla JM et al. A new titanosauriform sauropod from the Arcillas de Morella Formation (Early Cretaeous). In: Torcida Fernandez-Baldor F, Canudo JI, Huerta P, Pereda X (eds), Abstract Book of the VII International Symposium about Dinosaurs Palaeontology and their Environment. Salas de los Infantes: Colectivo Arqueologico y Paleontologico de Salas, 2016, 97-8.
  • Mocho P, Royo-Torres R, Ortega F. New data of the Portuguese brachiosaurid Lusotitan atalaiensis (Sobral Formation, Upper Jurassic). Historical Biology 2017a;29:789-817.
  • Mocho P, Perez-Garcia A, Gasulla JM et al. High sauropod diversity in the upper Barremian Arcillas de Morella Formation (Maestrat Basin, Spain) revealed by a systematic review of historical material.Journal of Iberian Geology 2017b;43:111-28.
  • Mocho P, Royo-Torres R, Escaso F et al. Upper Jurassic sauropod record in the Lusitanian Basin (Portugal): geographical and lithostratigraphical distribution. Palaeontologia Electronica 2017c;20:1-50, 27A.
  • Mocho P, Royo-Torres R, Ortega F. A new macronarian sauropod from the Upper Jurassic of Portugal. Journal of Vertebrate Paleontology 2019a;39:e1578782.
  • Mocho P, Perez-Garcia A, Martin Jimenez M et al. New remains from the Spanish Cenomanian shed light on the Gondwanan origin of European Early Cretaceous titanosaurs. Cretaceous Research 2019b;95:164-90.
  • Mocho P, Escaso F, Gasulla JM et al. New information about the sauropod dinosaurs of the Sant Antoni de la Vespa quarry (Morella, Spain). In: Blanco F, Blanco-Moreno C, Buscalioni AD, de la Cita L, Llandres M, Martin-Abad H, Marugan-Lobon J, Monleon MA, Navalon G, Nebreda SM, Prieto I, San Roman C (eds), Libro de Resumenes de las XXXVII Jornadas SEP y V Congreso Iberico de Paleontologia. Cuenca: Palaeontological publications 2, 2022, 116-116.
  • Moore AJ, Upchurch P, Barrett PM et al. Osteology of Klamelisaurus gobiensis (Dinosauria, Eusauropoda) and the evolutionary history of Middle-Late Jurassic Chinese sauropods. Journal of Systematic Palaeontology 2020;18:1299-393.
  • Moratalla JJ. Sauropod tracks of the Cameros Basin (Spain): identification, trackway patterns and changes over the Jurassic-Cretaceous. Geobios 2009;42:797-811.
  • Moratalla JJ, Jimenez S, Hernan S. Los Cayos dinosaur tracksite: an overview on the Lower Cretaceous ichno-diversity of the Cameros Basin (Cornago, La Rioja Province, Spain). Ichnos 2003;10:229-40.
  • O'Reilly JE, Puttick MN, Parry L et al. Bayesian methods outperform parsimony but at the expense of precision in the estimation of phylogeny from discrete morphological data. Biology Letters 2016;12:20160081.
  • Ortega F, Escaso F, Gasulla JM et al. Los Dinosaurios de la Peninsula Iberica. Estudios Geologicos 2006;62:219-40.
  • Osborn HF, Mook CC. Camarasaurus, Amphicoelias, and other sauropods of Cope.Memoirs of the American Museum of Natural History New Series 1921;3:247-387.
  • Ostrom JH, McIntosh JS. Marsh's Dinosaurs. New Haven: Yale University Press, 1966.
  • Otero A. The appendicular skeleton of Neuquensaurus, a Late Cretaceous saltasaurine sauropod from Patagonia, Argentina. Acta Palaeontologica Polonica 2010;55:399-426.
  • Otero A, Vizcaino SF. Hindlimb musculature and function of Neuquensaurus australis Lydekker (Sauropoda: Titanosauria). Ameghiniana 2008;45:333-48.
  • Otero A, Carballido JL, Moreno AP. The appendicular osteology of Patagotitan mayorum (Dinosauria, Sauropoda). Journal of Vertebrate Paleontology 2020;40:e1793158-19.
  • Paramo A, Mocho P, Ortega F. Three-dimensional analysis of the titanosaurian limb skeleton: implications for systematic analysis. Journal of Iberian Geology 2020;46:369-402.
  • Paramo A, Escaso F, Mocho P et al. 3D geometric morphometrics of the hind limb in the titanosaur sauropods from Lo Hueco (Cuenca, Spain). Cretaceous Research 2022;134:105147.
  • Pereda Suberbiola X, Ruiz-Omenaca JI. Los primeros descubrimientos de dinosaurios en Espana. Revista Espanola de Paleontologia 2005;10:15-28.
  • Pereda Suberbiola X, Torcida F, Izquierdo LA et al. First rebbachisaurid dinosaur (Sauropoda, Diplodocoidea) from the Early Cretaceous of Spain: palaeobiogeographical implications. Bulletin de la Societe Geologique de France 2003;174:471-9.
  • Perez-Garcia A, Sanchez Chillon B, Ortega F. Aportaciones de Jose Royo y Gomez al conocimiento sobre los dinosaurios de Espana. Paleolusitana 2009;1:339-64.
  • Perez Moreno A, Otero A, Carballido JL et al. The appendicular skeleton of Rinconsaurus caudamirus (Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina. Cretaceous Research 2023;142:105389.
  • Perez-Pueyo M, Moreno-Azanza M, Barco JL et al. New contributions to the phylogenetic position of the sauropod Galvesaurus herreroi from the late Kimmeridgian-early Tithonian (Jurassic) of Teruel (Spain). Boletin Geologico y Minero 2019;130:375-92.
  • Poropat SF,Upchurch P, Mannion PD et al. Revision of the sauropod dinosaur Diamantinasaurus matildae Hocknull et al. 2009 from the mid-Cretaceous of Australia:implications for Gondwanan titanosauriform dispersal. Gondwana Research 2015;27:995-1033.
  • Poropat SF, Mannion PD, Upchurch P et al. New Australian sauropods shed light on Cretaceous dinosaur palaeobiogeography. Scientific Reports 2016;6:34467.
  • Poropat SF, Mannion PD, Upchurch P et al. Osteology of the wide-hipped titanosaurian sauropod dinosaur Savannasaurus elliottorum from the Upper Cretaceous Winton Formation of Queensland, Australia. Journal of Vertebrate Paleontology 2020;40:e1786836.
  • Poropat SF, Kundrat M, Mannion PM et al. Second specimen of the Late Cretaceous Australian sauropod dinosaur Diamantinasaurus matildae provides new anatomical information on the skull and neck of early titanosaurs. Zoological Journal of the Linnean Society 2021;20:1-65.
  • Poropat SF, Frauenfelder TG, Mannion PD et al. Sauropod dinosaur teeth from the lower Upper Cretaceous Winton Formation of Queensland, Australia, and the global record of early titanosauriforms.Royal Society Open Science 2022;9:220381.
  • Poropat SF, Mannion PD, Rigby SL et al. A nearly complete skull of the sauropod dinosaur Diamantinasaurus matildae from the Upper Cretaceous Winton Formation of Australia and implications for the early evolution of titanosaurs. Royal Society Open Science 2023;10:221618.
  • PowellJE.Osteologiade Saltasaurusloricatus (Sauropoda-Titanosauridae) del Cretacico Superior del Noroeste argentino.In: Sanz JL, Buscalioni AD (eds), Los Dinosaurios y su Entorno Biotico. Cuenca: Instituto Juan de Valdes, Serie Actas Academicas, 1992, 165-230.
  • Poza B, Santos-Cubedo A, Santisteban C et al. Teredolites Leymerie 1842 from site with dinosaurs ANA (Lower Cretaceous, Cinctorres, Castellon). Geogaceta 2010;49:27-30.
  • Remes K. Revision of the Tendaguru sauropod Tornieria africana (Fraas) and its relevance for sauropod paleobiogeography.Journal of Vertebrate Paleontology 2006;26:651-69.
  • Riggs ES. Brachiosaurus altithorax, the largest known dinosaur. American Journal of Science 1903;s4-15:299-306.
  • Rose PJ. A new titanosauriform sauropod (Dinosauria: Saurischia) from the Early Cretaceous of central Texas and its phylogenetic relationships. Palaeontologica Electronica 2007;10:1-65.
  • Royo-Torres R. El sauropodo de Penarroya de Tastavins. Monograpfias Turonlenses 2009;6:1-548.
  • Royo-Torres R, Cobos A, Alcala L. A giant European dinosaur and a new sauropod clade. Science 2006;314:1925-7.
  • Royo-Torres R, Alcala L, Cobos A. A new specimen of the Cretaceous sauropod Tastavinsaurus sanzi from El Castellar (Teruel, Spain), and a phylogenetic analysis of the Laurasiformes. Cretaceous Research 2012;34:61-83.
  • Royo-Torres R, Upchurch P,Mannion PD et al.The anatomy,phylogenetic relationships and stratigraphic position of the Tithonian-Berriasian Spanish sauropod dinosaur Aragosaurus ischiaticus. Zoological Journal of the Linnean Society 2014;171:623-55.
  • Royo-Torres R, Fuentes C, Meijide M et al. A new Brachiosauridae Sauropod dinosaur from the lower Cretaceous of Europe (Soria Province, Spain). Cretaceous Research 2017a;80:38-55.
  • Royo-Torres R, Upchurch P, Kirkland JI et al. Descendants of the Jurassic turiasaurs from Iberia found refuge in the Early Cretaceous of western USA. Scientific Reports 2017b;7:14311.
  • Royo-Torres R, Cobos A, Mocho P et al. Origin and evolution of turiasaur dinosaurs set by means of a new 'rosetta' specimen from Spain. Zoological Journal of the Linnean Society 2021;191:201-27.
  • Royo y Gomez J.Los vertebrados del Cretacico espanol de facies wealdica. Boletin del Instituto Geologico y Minero de Espana 1926;47:171-6.
  • Ruiz-Omenaca JI, Pereda-Suberbiola X. Un documento inedito de Royo y Gomez sobre los dinosaurios del Levante. Temas Geologico-Mineros 1999;26:111-2.
  • Ruiz-Omenaca JI, Pereda Suberbiola X, Company J. Juan Vilanova, Jose Royo y Gomez y los dinosaurios de Morella en las colecciones del Museo Nacional de Ciencias Naturales de Madrid. In: XIX Jornadas de Paleontologia. Morella: Sociedade Espanola de Paleontologia, 2003, 150-1.
  • Salas R. El Malm i el Cretaci inferior entre el Massis de Garraf i la Serra d'Espada. Analisi de conca. Ph.D. Thesis, Universitat de Barcelona, 1987.
  • Salas R, Casas A. Mesozoic extensional tectonics, stratigraphy and crustal evolution during the Alpine cycle of the eastern Iberian basin. Tectonophysics 1993;228:33-55.
  • Salas R, Colombo F, Gamez D et al. Excursion guide book. In: XIX Jornadas de la Sociedad Espanola de Paleontologia. Morella: Sociedade Espanola de Paleontologia, 2003.
  • Salas R, Guimera J, Bover-Arnal T et al. The Iberian-Catalan Linkage: The Maestrat and Garraf Basins. In: Quesada C, Oliveira JT (eds), The Geology of Iberia: A Geodynamic Approach. Regional Geology Reviews, Vol.3. Berlin: Springer, 2020, 228-30.
  • Salgado L, Carvalho IS. Uberabatitan ribeiroi, a new titanosaur from the Marilia Formation (Bauru group, Upper Cretaceous), Minas Gerais, Brazil. Palaeontology 2008;51:881-901.
  • Salgado L, Coria RA, Calvo JO. Evolution of titanosaurid sauropods. I: phylogenetic analysis based on the postcranial evidence. Ameghiniana 1997;34:332.
  • Sanchez-Hernandez B, Benton MJ, Naish D. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. Palaegeography, Palaeoclimatology, Palaeoecology 2007;249:180-215.
  • Sander MP, Mateus O, Laven T et al. Bone histology indicates insular dwarfism in a new Late Jurassic sauropod dinosaur. Nature 2006;441:739-41.
  • Santisteban C, Santos-Cubedo A. The geological context: the Upper Jurassic and Lower Cretaceous. In: Galobart A, Suner M, Poza B (eds), Dinosaurs of Eastern Iberia (Life of the Past). Indiana: Indiana University Press, 2011, 40-55.
  • Santisteban C, Santos-Cubedo A, Poza B. Revision of the Mas de la Chimenea Alta fossil site section (Formacion Arcillas de Morella, Castellon) and its significance in the paleoenvironmental context. Geogaceta 2012;51:67-70.
  • Santisteban C, Santos-Cubedo A,Poza B et al. A different sedimentological look of the Arcillas de Morella Formation (Early Cretaceous, Eastern Iberia, Spain). In: Torcida Fernandez-Baldor F, Canudo JI, Huerta P, Pereda X (eds), Abstract Book of the VII International Symposium about Dinosaurs Palaeontology and their Environment. Salas de los Infantes: Colectivo Arqueologico y Paleontologico de Salas, 2016, 123-5.
  • Santos VF. Pegadas de Dinossaurios de Portugal. Lisboa: Museu Nacional de Historia Natural da Universidade de Lisboa, 2008.
  • Santos VF, Moratalla JJ, Royo-Torres R. New sauropod trackways from the Middle Jurassic of Portugal. Acta Palaeontologica Polonica 2009;54:409-22.
  • Santos-Cubedo A, de Santisteban C, Galobart A. New dinosaur findings from Arcillas de Morella Formation (Spain).In: Godefroit P, Lambert O (eds), Tribute to Charles Darwin and Bernissart Iguanodons: New Perspectives on Vertebrate Evolution and Early Cretaceous Ecosystems. Brussels, 2009, 87.
  • Santos-Cubedo A, Poza B, Suner M et al. New remains of a titanosaur (Dinosauria: Sauropoda) from the Early Cretaceous of Spain. Journal of Vertebrate Paleontology Supplement to Number 2010;30:157A.
  • Santos-Cubedo A, de Santisteban C, Poza B et al. Using sequence stratigraphy to understand the taphonomy of dinosaur sites in the Arcillas de Morella Formation (Lower Cretaceous, Eastern Iberia,Spain).In: The Geological Society (ed.), Abstract Book of the William Smith Meeting 2014: The Future of Sequence Stratigraphy: Evolution or Revolution? London: The Geological Society, 2014, 77-77.
  • Santos-Cubedo A, Santisteban C. de, Poza B et al. A new spinosaurid dinosaur species from the Early Cretaceous of Cinctorres (Spain). Scientific Reports 2023;13:6471.
  • Sanz JL. Las faunas espanolas de dinosaurios. In: I Congreso Espanol de Geologia, Segovia. Espanha: Ilustre Colegio Oficial de Geologos, 1984, 497-506.
  • Sanz JL. Jose Royo y Gomez y los dinosaurios espanoles. Geogaceta 1996;19:167-8.
  • Sanz JL, Casanovas ML, Santafe JV. II. Paleontologia. In: Santafe JV, Casanovas ML, Sanz JL, Calzada S (coord.), Geologia y Paleontologia (Dinosaurios) de las Capas rojas de Morella (Castellon, Espana). Barcelona: Diputacion Provincial de Castellon y Diputacion de Barcelona, 1982, 71-169.
  • Sauvage HE. Vertebres fossiles du Portugal. Contribution a l'etude des Poissons et des Reptiles du Jurassique et du Cretace. Lisboa: Memoires de la Direction des Travaux Geologiques du Portugal, 1897-98.
  • Sereno PC, Wilson JA, Witmer LM et al. Structural extremes in a Cretaceous dinosaur. PLoS One 2007;2:e1230.
  • Silva Junior JCG, Marinho TS, Martinelli AG et al. Osteology and systematics of Uberabatitan ribeiroi (Dinosauria; Sauropoda): a Late Cretaceous titanosaur from Minas Gerais, Brazil. Zootaxa 2019;4577:401-38.
  • Suteethorn S, Le Loeuff J, Buffetaut E et al. A new skeleton of Phuwiangosaurus sirindhornae (Dinosauria, Sauropoda) from NE Thailand. Geological Society, London, Special Publications 2009;315:189-215.
  • Sweetman SC. A gobiconodontid (Mammalia, Eutriconodonta) from the Early Cretaceous (Barremian) Wessex Formation of the Isle of Wight, southern Britain. Palaeontology 2006;49:889-97.
  • Sweetman SC. A spalacolestine spalacotheriid (Mammalia, Trechnotheria) from the Early Cretaceous (Barremian) of southern England and its bearing on spalacotheriid evolution. Palaeontology 2008;51:1367-85.
  • Tidwell V, Carpenter K, Brooks W. New sauropod from the Lower Cretaceous of Utah, USA. Oryctos 1999;2:21-37.
  • Tidwell V, Carpenter K, Meyer S. New titanosauriform (Sauropoda) from the Poison Strip Member of the Cedar Mountain Formation (Lower Cretaceous), Utah. In: Tanke DH, Carpenter K (eds), Mesozoic Vertebrate Life. Bloominton and Indianapolis: Indiana University Press, 2001, 139-65.
  • Torcida Fernandez-Baldor F. Sistematica, filogenia y analisis paleobiogeografico de Demandasaurus darwini (Sauropoda, Rebbachisauridae) del Barremiense Superior-Aptiense de Burgos (Espana). Unpublished Ph.D. Thesis, Universidad Zaragoza, 2012
  • Torcida Fernandez-Baldor FT, Canudo JI, Huerta P et al. Demandasaurus darwini, a new rebbachisaurid sauropod from the Early Cretaceous of the Iberian Peninsula. Acta Palaeontologica Polonica 2011;56:535-52.
  • Torcida Fernandez-Baldor F, Canudo JI, Huerta P et al. Europatitan eastwoodi, a new sauropod from the Lower Cretaceous of Iberia in the initial radiation of somphospondylans in Laurasia. PeerJ 2017;5:e3409.
  • Tschopp E, Mateus O. Clavicles, interclavicles, gastralia, and sternal ribs in sauropod dinosaurs: new reports from Diplodocidae and their morphological, functional and evolutionary implications. Journal of Anatomy 2013;222:321-40.
  • Tschopp E, Upchurch P. The challenges and potential utility of phenotypic specimen-level phylogeny based on maximum parsimony. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 2019;109:301-23.
  • Tschopp E, Mateus O, Benson RBJ. A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ 2015;3:e857.
  • Upchurch P.The evolutionary history of sauropod dinosaurs.Philosophical Transactions of the Royal Society of London, Series B 1995;349:365-90.
  • Upchurch P. The phylogenetic relationships of sauropod dinosaurs. Zoological Journal of the Linnean Society 1998;124:43-103.
  • Upchurch P, Martin J. The anatomy and taxonomy of Cetiosaurus (Saurischia: Sauropoda) from the Middle Jurassic of England. Journal of Vertebrate Paleontology 2003;23:208-31.
  • Upchurch P, Barrett PM, Dodson P. Sauropoda. In: Weishampel DB, Dodson P, Osmolska H (eds), The Dinosauria II. Berkeley: University of California, 2004, 259-322.
  • Upchurch P, Mannion PD, Barrett PM. Sauropod dinosaurs. In: Batten DJ (ed.), English Wealden Fossils, Vol. 14. London: Palaeontological Association, 2011, 476-525.
  • Upchurch P, Mannion PD, Taylor MP. The Anatomy and phylogenetic relationships of "Pelorosaurus" becklesii (Neosauropoda, Macronaria) from the Early Cretaceous of England. PLoS One 2015;10:e0125819.
  • Vila B, Galobart A, Canudo JI et al. The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe:clues to demise and extinction.Palaeogeography, Palaeoclimatology, Palaeoecology 2012;350-352:19-38.
  • Vilanova y Piera J.Compendio de Geologia. Madrid:Imprenta de Alejandro Gomez Fuentenebro, 1872.
  • Vilas L, Mas R, Garcia A et al. Capitulo 8. Iberica Suroccidental. In: Dept. de Estratigrafia de la Fac. de Ciencias Geologicas de la Universidad Complutense de Madrid, Unidad Estructuralde investigacion de Correlaciones estratigraficas y paleogeografia del Instituto de Geologia Economica (eds), El Cretacico de Espana. Madrid: Universidad Complutense, Madrid, 1982, 457-509.
  • Villanueva-Amadoz U, Santisteban C, Santos-Cubedo A. Age determination of the Arcillas de Morella Formation (Maestrazgo Basin, Spain). Historical Biology 2014;27:389-97.
  • Vullo R, Marugan-Lobon J, Kellner AWA et al. A new crested pterosaur from the Early Cretaceous of Spain: The first European tapejarid (Pterodactyloidea: Azhdarchoidea). PLoS One 2012;7:e38900.
  • Wang X, You H, Meng Q et al. Dongbeititan dongi, the first sauropod dinosaur from the Lower Cretaceous Jehol Group of western Liaoning Province, China. Acta Geologica Sinica 2007;81:911-6.
  • Wedel MJ. The evolution of vertebral pneumaticity in sauropod dinosaurs. Journal of Vertebrate Paleontology 2003;23:344-57.
  • Wedel MJ, Cifelli RL, Sanders RK. Osteology, paleobiology, and relationships of the sauropod dinosaur Sauroposeidon. Acta Palaeontologica Polonica 2000;45:343-88.
  • Whitlock JA. A phylogenetic analysis of Diplodocoidea (Saurischia: Sauropoda). Zoological Journal of the Linnean Society 2011;161:872-915.
  • Wilson JA. A nomenclature for vertebral laminae in sauropods and other saurischian dinosaurs. Journal of Vertebrate Paleontology 1999;19:639-53.
  • Wilson JA. Sauropod dinosaur phylogeny: critique and cladistic analysis. Zoological Journal of the Linnean Society 2002;136:215-75.
  • Wilson JA. Anatomical nomenclature of fossil vertebrates: standardized terms or lingua franca? Journal of Vertebrate Paleontology 2006;26:511-8.
  • Wilson JA.New vertebral laminae and patterns of serial variation in vertebral laminae of sauropod dinosaurs. Contributions from the Museum of Paleontology 2012;32:91-110.
  • Wilson JA, Allain R. Osteology of Rebbachisaurus garasbae Lavocat, 1954, a diplodocoid (Dinosauria, Sauropoda) from the early Late Cretaceous-aged Kem Kem beds of southeastern Morocco. Journal of Vertebrate Paleontology 2015;35:e1000701.
  • Wilson JA, Carrano MT. Titanosaurs and the origin of 'wide-gauge' trackways: a biomechanical and systematic perspective on sauropod locomotion. Paleobiology 1999;25:252-67.
  • Wilson JA, Sereno PC. Early evolution and higher-level phylogeny of sauropod dinosaurs. Journal of Vertebrate Paleontology 1998;5:1-68.
  • Wilson JA, Upchurch P. Redescription and reassessment of the phylogenetic affinities of Euhelopus zdanskyi (Dinosauria: Sauropoda) from the Early Cretaceous of China. Journal of Systematic Palaeontology 2009;7:199-239.
  • Wilson JA, D'Emic MD, Ikejiri T et al. A nomenclature for vertebral fossae in sauropods and other saurischian dinosaurs. PLoS One 2011a;6:e17114.
  • Wilson JA, Barrett PM, Carrano MT. An associated partial skeleton of Jainosaurus cf. septentrionalis (Dinosauria: Sauropoda) from the Late Cretaceous of Chhota Simla, Central India. Palaeontology 2011b;54:981-98.
  • Wiman C. Die Kreide-Dinosaurier aus Shantung. Palaeontologia Sinica (Series C) 1929;6:1-67.
  • Xu X, Upchurch P, Mannion PD et al. A new Middle Jurassic diplodocoid suggests an earlier dispersal and diversification of sauropod dinosaurs. Nature Communications 2018;9:2300.
  • Yague P, Upchurch P, Sanz JL et al. New sauropod material from the Early Cretaceous of Spain. In: 49 Annual Symposium of Vertebrate Paleontology and Comparative Anatomy, Yorkshire Museum, 2001.
  • You H-L, Tang F, Luo Z. A new basal titanosaur (Dinosauria: Sauropoda) from the Early Cretaceous of China. Acta Geologica Sinica 2003;77:424-9.
  • You H-L, Li D-Q, Zhou L-Q, et al. Daxiatitan binglingi: a giant sauropod dinosaur from the Early Cretaceousof China. Gansu Geology 2008:17:1-10.
  • Zaher H, Pol D, Carvalho AB et al. A complete skull of an Early Cretaceous sauropod and the evolution of advanced titanosaurians. PLoS One 2011;6:e16663.