Published October 24, 2023 | Version v1
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Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae)

  • 1. State Museum of Natural History Stuttgart, Entomology, Rosenstein & D-70191 Stuttgart, Germany. & University of Hohenheim, Systematic Entomology (190n), Garbenstr. 30, D-70599 Stuttgart, Germany.
  • 2. SNSB-ZSM, Bavarian State Collection of Zoology, Münchhausenstr. 21, D-81247 Munich, Germany.
  • 3. Finnish Museum of Natural History, Zoology Unit, University of Helsinki, P.O. Box
  • 4. Grupo Biología Evolutiva, Department of Biology, Universidad de Sucre, Sincelejo, Sucre, Colombia. Corresponding author.
  • 5. State Museum of Natural History Stuttgart, Entomology, Rosenstein & D-70191 Stuttgart, Germany.

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Wanke, Dominic, Hausmann, Axel, Lee, Kyung Min, Murillo-Ramos, Leidys, Sihvonen, Pasi, Rajaei, Hossein (2023): Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae). Zootaxa 5359 (1): 1-96, DOI: 10.11646/zootaxa.5359.1.1, URL: https://www.mapress.com/zt/article/download/zootaxa.5359.1.1/52094

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References

  • Ban, X., Jiang, N., Cheng, R., Xue, D. & Han, H. (2018) Tribal classification and phylogeny of Geometrinae (Lepidoptera: Geometridae) inferred from seven gene regions. Zoological Journal of the linnean Society, 184 (3), 653-672. https://doi.org/10.1093/zoolinnean/zly013
  • Barou, P.J. (1967) Contribution a la connaissance la faune des Lepidopteres de l'Iran. Entomologie et Phytopathologie Appliquees, 26, 41-58.
  • Beljaev, E.A. (2016) Fam. Geometridae-Pyadenitzi. In: Lelei, A.S. (Ed.), Annotated catalogue of the insects of Russian Far East. Vol. 2. Lepidoptera. Dal'nauka, Vladivostok, pp. 518-666. [in Russian]
  • Bienert, Th. (1869) Lepidopterologische Ergebnisse einer Reise in Persien in den Jahren 1858 und 1859. Inaugural-Dissertation zur Erlangung der philosophischen Doctorwurde, an der Universitat Leipzig, Leipzig, 56 pp.
  • Brandt, W. (1938) Beitrag zur Lepidopteren-Fauna von Iran. Neue Gattungen, Arten und Formen (Macrolepidoptera). Entomologische Rundschau, 55 (49), 567-574.
  • Brandt, W. (1939) Beitrag zur Lepidopteren-Fauna von Iran. Neue Gattungen, Arten und Formen (Macrolepidoptera). Entomologische Rundschau, 56 (14), 139-142.
  • Brandt, W. (1941) Beitrag zur Lepidopteren-Fauna von Iran (4). Einige neue Geometriden. Mitteilungen der Munchner Entomologischen Gesellschaft, 31 (3), 864-886.
  • Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B.C., Ounap, E., Moser, A., Mˆ rtter, R., Bolt, D., Bodner, F., Lindt, A., Parra, L.E. & Wahlberg, N. (2019) New World geometrid moths (Lepidoptera: Geometridae): molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Systematics & Phylogeny, 77 (3), 457-486. https://doi.org/10.26049/ASP77-3-2019-5
  • Choi, S.W. & Kim, S.S. (2016) A checklist of the genus Scopula (Lepidoptera: Geometridae) including description of a new species and three newly recorded species from Korea. Zootaxa, 4178 (1), 131-137. https://doi.org/10.11646/zootaxa.4178.1.6
  • Christoph, H.T. (1873) Weiterer Beitrag zum Verzeichnisse der in Nord-Persien einheimischen Schmetterlinge. Horae Societatis Entomologicae Rossicae, 10, 3-55.
  • Christoph, H.T. (1876-1877) Sammelergebnisse aus Nordpersien, Krasnowodsk in Turkmenien und dem Daghestan. Horae Societatis Entomologicae Rossicae, 12, 181-299.
  • Covell, C.V. (1970) A revision of the North American species of the genus Scopula (Lepidoptera: Geometridae). Transactions of the American Entomological Society, 96, 101-221.
  • Eskandarzadeh, N., Rastegar-Pouyani, N., Rastegar-Pouyani, E., Fathinia, B., Bahmani, Z., Hamidi, K. & Gholamifard, A. (2018) Annotated checklist of the endemic Tetrapoda species of Iran. Zoosystema, 40 (sp1), 507-537. https://doi.org/10.5252/zoosystema2018v40a24
  • Feizpour, S., Fekrat, L., Namaghi, H.S., Stadie, D. & Rajaei, H. (2018) Combination of morphological characters and DNAbarcoding confirms Problepsis cinerea (Butler, 1886) (Geometridae: Sterrhinae: Scopulini) as a new genus and species for the fauna of Iran. Integrative Systematics, 1, 47-57. https://doi.org/10.18476/insy.v01.a6
  • Gholamifard, A. (2011) Endemism in the reptile fauna of Iran. Iranian Journal of Animal Biosystematics, 7 (1), 13-29.
  • Hausmann, A. (1993) Zweiter Beitrag zur Taxonomie und Systematik der Gattung Glossotrophia Prout, 1913 (Lepidoptera, Geometridae, Sterrhinae). Mitteilungen der Munchener Entomologischen Gesellschaft, 83, 77-107.
  • Hausmann, A. (1994) Dritter Beitrag zur Revision der Gattung Glossotrophia Prout, 1913 nebst Beschreibung zweier neuer Gattungen (Lepidoptera: Geometridae, Sterrhinae). Nota Lepidopterologica, 16, 195-211.
  • Hausmann, A. (1996) Zwei neue Geometridenarten aus Vorderasien (Insecta, Lepidoptera, Geometridae). Spixiana Supplement, 22, 3-10.
  • Hausmann, A. (2004) The Geometrid Moths of Europe. Vol. 2. Sterrhinae. Apollo Books, Stenstrup, 600 pp. https://doi.org/10.1163/9789004322554
  • Hausmann, A. & Dˆ tterl, S. (2004) Nectar-plants and larval food-plants of the genus Glossotrophia (Lepidoptera: Geometridae, Sterrhinae): studies on pollen grains attached to museum specimens. Nota Lepidopterologica, 26 (3/4), 127-136.
  • Hausmann,A. & Laszlo G.M. (1999) Taxonomic and faunistic studies on Turkmenian Sterrhinae. Folia Entomologica Hungarica, 60, 317-324.
  • Hausmann, A., Miller, S.E., Holloway, J.D., deWaard, J., Pollock, D., Prosser, S.W.J. & Hebert, P.D.N. (2016a) Calibrating the taxonomy of a megadiverse family on BOLD: 3000 DNA barcodes from geometrid type specimens (Lepidoptera, Geometridae). Genome, 59 (9), 671-684. https://doi.org/10.1139/gen-2015-0197
  • Hausmann, A., Stadie, D. & Fiebig, R. (2016b) Family Geometridae Leach, [1815]. In: Hacker, H. (Ed.), Systematic and illustrated catalogue of the Macroheterocera and superfamilies Cossoidea Leach, [1815], Zygaenoidea Latreille, 1809, Thyridoidea Herrich-Schaffer, 1846 and Hyblaeoidea Hampson, 1903 of the Arabian Peninsula, with a survey of their distribution (Lepidoptera). Esperiana, 20 (1), pp. 61-138.
  • Hausmann, A., Kravchenko, V. & Muller, G. (2020) The Lepidoptera of Israel. Vol. 3. Geometridae. Proceedings of the Museum Witt Munich, 9, 1-256.
  • Hufnagel, J.S. (1767) Fortsetzung der Tabelle von den Nachtvoegeln, welche die 3te Art derselben, nehmlich die Spannenmesser (Phalaenas Geometras Linnaei) enthalt. Berlinisches Magazin, 4 (5), 504-527.
  • Ivanova, N.V., deWaard, J.R. & Hebert, P.D.N. (2006) An inexpensive, automation-friendly protocol for recovering high-quality DNA. Molecular Ecology Notes, 6, 998-1002. https://doi.org/10.1111/j.1471-8286.2006.01428.x
  • Jiang, N., Li, X.X., Hausmann, A., Cheng, R., Xue, D.Y. & Han, H.X. (2017) A molecular phylogeny of the Palaearctic and Oriental members of the tribe Boarmiini (Lepidoptera: Geometridae: Ennominae). Invertebrate Systematics, 31 (4), 427- 441. https://doi.org/10.1071/IS17005
  • Kalyaanamoorthy, S., Minh, B.Q., Wong, T.K.F., von Haeseler, A. & Jermiin, L.S. (2017) ModelFinder: Fast model selection for accurate phylogenetic estimates. Nature Methods, 14, 587-589. https://doi.org/10.1038/nmeth.4285
  • Kalali, Gh.-H. (1976) A List of Lepidoptera from Province of Khorasan (Iran). Journal of Entomological Society of Iran, 3, 131-135.
  • Kimura, M. (1980) A simple method for estimating evolutionary rates of base substitutions through comparative studies of nucleotide sequences. Journal of Molecular Evolution, 16, 111-120. https://doi.org/10.1007/BF01731581
  • Kouznetsov, V.J. (1959) Sur la Faune des Lepidopteres du Sud de l'Iran. Bulletin de la Societe Entomologique de Mulhouse, 15, 65-70.
  • Kumar, S., Stecher, G., Li, M., Knyaz, C. & Tamura, K. (2018) MEGA X: Molecular Evolutionary Genetics Analysis across computing platforms. Molecular Biology and Evolution, 35 (6), 1547-1549. https://doi.org/10.1093/molbev/msy096
  • Lederer, J. (1871) Nachtrag zum Verzeichnisse der von Herrn Jos. Haberhauer bei Astrabad in Persien gesammelten Schmetterlinge. Horae Societatis Entomologicae Rossicae, 8, 3-28.
  • Lehmann, L., Stadie, D. & Zahiri, R. (2009) Zum Winteraspekt der Makrolepidopterenfauna Sudirans mit Anmerkungen zur Biologie einiger Arten (Lepidoptera: Bombycoidea, Papilionoidea, Geometroidea, Noctuoidea). Nachrichten des Entomologischen Vereins Apollo, Neue Folge, 30 (3), 105-119.
  • Lehmann, L. & Zahiri, R. (2011) Results of a lepidopterological expedition to North and Northwest Iran in summer 2007 with new records for Iran (Lepidoptera). Esperiana, 16 135-165.
  • Makhov, I. (2023) Geometridae (Lepidoptera) of the Baikal region: identification keys and annotated catalogue with notes to DNA barcoding. Part 2. Archiearinae, Geometrinae, Sterrhinae. Zootaxa, 5294 (1), 1-120. https://doi.org/10.11646/zootaxa.5294.1.1
  • Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. Proceedings of the Gateway Computing Environments Workshop (GCE), New Orleans, Louisiana, 11705685. https://doi.org/10.1109/GCE.2010.5676129
  • Muller, B., Erlacher, S., Hausmann, A., Rajaei, H., Sihvonen, P. & Skou, P. (2019) Ennominae II. In: Hausmann, A., Rajaei, H., Shivonen, P. & Skou, P. (Eds.), The Geometrid Moths of Europe. Vol. 6. Brill, Leiden, pp. 1-906. https://doi.org/10.1163/9789004387485
  • Murillo-Ramos, L., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Reza Ghanavi, H., Ounap, E., Truuverk, A., Staude, H., Friedrich, E., Tammaru, T. & Wahlberg, N. (2019) A comprehensive molecular phylogeny of Geometridae (lepidoptera) with a focus on enigmatic small subfamilies. PeerJ, 7, e7386. https://doi.org/10.7717/peerj.7386
  • Noori, S., Wanke, D. & Rajaei, H. (2023) A gazetteer of elusive localities of Iranian Lepidoptera records. In: Rajaei, H. & Karsholt, O. (Eds.), Lepidoptera Iranica. Integrative Systematics, 6 (Special Issue), pp. 91-129. https://doi.org/10.18476/2023.997558.6
  • Noroozi, J., Talebi, A., Doostmohammadi, M., Manafzadeh, S., Asgarpour, Z. & Schneeweiss, G.M. (2019) Endemic Diversity and Distribution of the Iranian Vascular Flora across Phytogeographical Regions, Biodiversity Hotspots and Areas of Endemism. Scientific Reports, 9 (1), 1-12. https://doi.org/10.1038/s41598-019-49417-1
  • Ot unap, E. (2010) Systematic studies on the subfamily Sterrhinae (Lepidoptera: Geometridae). Academic Dissertation, University of Tartu, Tartu, 111 pp. [ISBN 978-9949-19-361-5]
  • Ounap, E., Viidalepp, J. & Truuverk, A. (2016) Phylogeny of the subfamily Larentiinae (Lepidoptera: Geometridae): integrating molecular data and traditional classifications. Systematic Entomology, 41, 824-843. https://doi.org/10.1111/syen.12195
  • Prosser, S.W.P., deWaard, J.R., Miller, S.E. & Hebert, P.D.N. (2016) DNA barcodes from century-old type specimens using next-generation sequencing. Molecular Ecology Resources, 16 (2), 487-497. https://doi.org/10.1111/1755-0998.12474
  • Prout, L.B. (1912-1916) Die spannerartigen Nachtfalter. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 4. A. Kernen verlag, Stuttgart, pp. 1-479.
  • Prout, L.B. (1921) Moths of Mesopotamia and N. W. Persia, Part III. Geometridae. Journal Bombay Natural History Society, XXVIII (1 & 2), 187-191.
  • Rajaei, H., Hausmann, A., Scoble, M., Wanke, D., Plotkin, D., Brehm, G., Murillo-Ramos, L. & Sihvonen, P. (2022): An online taxonomic facility of Geometridae (Lepidoptera), with an overview of global species richness and systematics. Integrative Systematics, 5 (2), 145-192. https://doi.org/10.18476/2022.577933
  • Rajaei, H., Aarvik, L., Arnscheid, W.R., Baldizzone, G., Bartsch, D., Bengtsson, B.A., Bidzilya, O., Buchner, P., Buchsbaum, U., Buszko, J., Dubatolov, V.V., Erlacher, S., Esfandiari, M., Freina, J.J. De, Gaedike, R., Gyulai, P., Hausmann, A., Haxaire, J., Hobern, D., Hofmann, A., Ignatev, N., Kaila, L., Kallies, A., Keil, T., Kiss, A., Kitching, I.J., Kun, A., Laszlo, G.M., Leraut, G., Mally, R., Matov, A., Meineke, J.-U., Melichar, T., Mey, W., Mironov, V., Muller, B., Naderi, A., Nassig, W.A., Naumann, S., Nazari, V., Nieukerken, E.J. Van, Nuss, M., Pˆ ll, N., Prozorov, A.M., Rabieh, M.M., Rakosy, L., Rindos, M., Rota, J., Rougerie, R., Schintlmeister, A., Shirvani, A., Sihvonen, P., Simonsen, T.J., Sinev, S.Yu., Skou, P., Sobczyk, T., Sohn, J.-C., Tabell, J., Tarmann, G., Tokar, Z., Trusch, R., Varga, Z., Volynkin, A.V., Wanke, D., Yakovlev, R.V., Zahiri, R., Zehzad, P., Zeller, H.C., Zolotuhin, V.V. & Karsholt, O. (2023a) Catalogue of the Lepidoptera of Iran. In: Rajaei, H. & Karsholt, O. (Eds.), Lepidoptera Iranica. Integrative Systematics, 6 (Special Issue), pp. 121-459. https://doi.org/10.18476/2023.997558.7
  • Rajaei, H., Noori, S., Karsholt, O. & Zahiri, R. (2023b) General patterns of the Lepidoptera fauna of Iran. In: Rajaei, H. & Karsholt, O. (Eds.), Lepidoptera Iranica. Integrative Systematics, 6 (Special Issue), pp. 69-81. https://doi.org/10.18476/2023.997558.4
  • Rajaei, H., Stuning, D. & Trusch, R. (2012) Taxonomic revision and zoogeographical patterns of the species of Gnopharmia Staudinger, 1892 (Geometridae, Ennominae). Zootaxa, 3360 (1), 1-52. https://doi.org/10.11646/zootaxa.3360.1.1
  • Rambaut, A. (2012) Figtree. Version 1.4.3. Available from: http://tree.bio.ed.ac.uk/software/figtree/ (accessed 6 September 2023)
  • Reisser, H. (1958) Ergebnisse der Osterreichischen Iran-Expedition 1949/50 Lepidoptera I. (Macrolepidoptera). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, Abtheilung 1, 167 (10), 519-551.
  • Romanoff, N.M. (1885) Les Lepidopteres de la Transcaucasie. Deuxieme partie. In: Romanoff, N.M. (Ed.), Memoires sur les Lepidopteres, 2, pp. 1-118.
  • Sannino, L. & Espinosa, B., (1999) Aspetti morfologici ed etologici di Scopula turbidaria (Lepidoptera: Geometridae). Fragmenta entomologica, Roma, 31 (2), 377-395.
  • Schwingenschuss, L. (1939) Beitrag zur Lepidopterenfauna von Iran (Persien) insbesondere des Elbursgebirges in Nordiran. (Fortsetzung). Entomologische Zeitschrift, 53, 200-210.
  • Scoble, M.J. (1999) Geometrid Moths of the World: A Catalogue (Lepidoptera, Geometridae). Vols. 1 & 2. CSIRO Publishing, Collingwood, Victoria and Apollo Books, Stenstrup, 1200 pp.
  • Scoble, M.J. & Hausmann, A. (2007) Online list of valid and available names of the Geometridae of the World. Updated. Available from: https://geometroidea.smns-bw.org/archive/ (accessed 19 Ferbruary 2023)
  • Sihvonen, P. (2001) Revision of the Scopula cajanderi (Lepidoptera: Geometridae: Sterrhinae) species group with description of a new species. The Canadian Entomologist, 133, 467-486. https://doi.org/10.4039/Ent133467-4
  • Sihvonen, P. (2003) Diversity and classification of the Scopulini moths (Lepidoptera: Geometridae, Sterrhinae). Academic Dissertation, University of Helsinki, Helsinki, 158 pp. [Edta Prima Oy, ISBN 952-10-1400-8]
  • Sihvonen, P. & Kaila, L. (2004) Phylogeny and tribal classification of Sterrhinae with emphasis on delimiting Scopulini. Systematic Entomology, 29, 324-358. https://doi.org/10.1111/j.0307-6970.2004.00248.x
  • Sihvonen, P. (2005a) Phylogeny and classification of the Scopulini moths (Lepidoptera: Geometridae, Sterrhinae). Zoological Journal of Linnean Society, 143, 473-530. https://doi.org/10.1111/j.1096-3642.2005.00153.x
  • Sihvonen, P. (2005b) Check-list of Chinese Scopula Schrank species and an analysis of species diversity (Lepidoptera: Geometridae: Sterrhinae). Journal of Asia-Pacific Entomology, 8, 29-36. https://doi.org/10.1016/S1226-8615(08)60069-2
  • Sihvonen, P., Murillo-Ramos, L., Brehm, G., Staude, H. & Wahlberg, N. (2020) Molecular phylogeny of Sterrhinae moths (Lepidoptera: Geometridae): towards a global classification. Systematic Entomology, 45, 606-634. https://doi.org/10.1111/syen.12418
  • Stadie, D. & Stadie, Ph. (2016): The Problepsis ocellata Frivaldszky, 1845 species lineage - an integrative survey of their morphology, species radiation, distribution and ecology (Lepi- doptera, Geometridae, Sterrhinae). Esperiana, 20 (2), 381- 429.
  • Stamatakis, A. (2014) RAxML Version 8: A tool for phylogenetic analysis and post-analysis of large phylogenies. Bioinformatics, 30, 1312-1313. https://doi.org/10.1093/bioinformatics/btu033
  • Stamatakis, A., Hoover, P. & Rougemont, J. (2008) A rapid bootstrap algorithm for the RAxML web-servers. Systematic Biology, 75, 758-771. https://doi.org/10.1080/10635150802429642
  • Sobczyk, T. & Gelbrecht, J. (2004) Zur Arealregression der an Thymian (Thymus) gebundenen Arten Scopula decorata ([Denis & Schiffermuller], 1775) und Pseudophilotes vicrama (Moore, 1864) in Deutschland (Lepidoptera, Geometridae et Lycaenidae). Markische Entomologischen Nachrichten, 6 (1), 1-16.
  • Sutton, S.L. (1963) South Caspian Insect Fauna 1961. I. Systematic List of Lepidoptera with notes. The Annals and magazine of natural history; zoology, botany, and geology, 13 (6), 353-374. https://doi.org/10.1080/00222936308651370
  • Viidalepp, J. (1988) Geometridae fauna of the Central Asian mountains. Nauka Moscow, 240 pp.
  • Viidalepp, J. (1996) Checklist of the Geometridae (Lepidoptera) of the former U.S.S.R. Apollo Books, Stenstrup, 111 pp.
  • Wahlberg, N., Pena, C., Ahola, M., Wheat, C.W. & Rota J. (2016) PCR primers for 30 novel gene regions in the nuclear genomes of lepidoptera. Zookeys, 596, 129-141. https://doi.org/10.3897/zookeys.596.8399
  • Wahlberg, N. & Wheat, C.W. (2008) Genomic outposts serve the phylogenomic pioneers: designing novel nuclear markers for genomic DNA extractions of Lepidoptera. Systematic Biology, 57, 231-242. https://doi.org/10.1080/10635150802033006
  • Wanke, D., Bigalk, S., Krogmann, L., Wendt, I. & Rajaei, H. (2019) The Fixator-A simple method for mounting of arthropod specimens and photography of complex struc- tures in liquid. Zootaxa, 4657 (2), 385-391. https://doi.org/10.11646/zootaxa.4657.2.11
  • Wanke, D., Hausmann, A., Krogmann, L., Petranyi, G. & Rajaei, H. (2020) Taxonomic revision of the genus Nychiodes Lederer, 1853 (Geometridae: Ennominae: Boarmiini) with description of three new species-an integrative approach. Zootaxa, 4812 (1), 1-61. https://doi.org/10.11646/zootaxa.4812.1.1
  • Wanke, D., Krogmann, L., Murillo-Ramos, L., Sihvonen, P. & Rajaei, H. (2021b) Systematics of Problepsis wiltshirei (Prout, 1938), comb. nov. (Lepidoptera, Geometridae, Sterrhinae)-an endemic species to the Zagros Mountains in the Middle East. Nota Lepidopterologica, 44, 175-192. https://doi.org/10.3897/nl.44.67345
  • Wanke, D. & Rajaei, H. (2018) An effective method for the close up photography of insect genitalia during dissection: a case study on the Lepidoptera. Nota Lepidopterologica, 41 (2), 219-223. https://doi.org/10.3897/nl.41.27831
  • Wanke D., Ulmer J.M., Wendt I. & Rajaei H. (2021a) Updates on the Fixator-Facilitating the investigation, mounting, and photography of structures and specimens in liquid. Zootaxa, 4999 (4), 397-400. https://doi.org/10.11646/zootaxa.4999.4.9
  • Wehrli, E. (1939-1954) Die Spanner des Palaearktischen Faunengebietes. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 4. Supplement. A. Kernen, Stuttgart, pp. 254-766.
  • Werner, M.J., Hausmann, A., Kostjuk, I., Wanke, D. & Rajaei, H. (2023) Integrative taxonomic revision of the genus Phaselia Guenee, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia. Zootaxa, 5326 (1), 1-66. https://doi.org/10.11646/zootaxa.5326.1.1
  • Wieser, Ch., Huemer, P. & Stangelmaier, G. (2002) In: Gutleb, B. & Wieser, Ch. (2002) Ergebnisse einer zoologischen Exkursion in den Nordiran, 2001. Carinthia II, 192/112, pp. 33-140.
  • Wieser, Ch. & Stangelmaier, G. (2005) Zwischenergebnisse einer lepidopterologischen Forschungsreise in den Nordiran, Oktober 2003 (Insecta: Lepidoptera). Carinthia II, 195/115, 659-674.
  • Wiltshire, E.P. (1941) Mesopotamian Desert Lepidoptera. The journal of the Bombay Natural History Society, 42 (4), 826- 842.
  • Wiltshire, E.P. (1943) Early stages of oriental palearctic Lepidoptera. V. Journal of the Bombay Natural History Society, Bombay, 43, 621-634.
  • Wiltshire, E.P. (1948) Middle East Lepidoptera IX: Two new forms or species and thirty-five new records from Cyprus. The Entomologist's Record and Journal of Variation, 60, 79-87.
  • Wiltshire, E.P. (1957) The Lepidoptera of Iraq. Ministry of Agriculture, Government of Iraq, Baghdad, 162 pp.
  • Wiltshire, E.P. (1966) Osterreichische entomologische Iran-Afghanistan-Expeditionen, Beitrage zur Lepidopterenfauna, Teil 9, Subfamilie Sterrhinae (Lepidoptera, Geometridae). Middle East Lepidoptera XXII. Zeitschrift der Wiener Entomologischen Gesellschaft, 51 (9-11), 113-138.
  • Wiltshire, E.P. (1967) Middle East Lepidoptera XX.A third contribution to the Fauna of Afghanistan.Beitrage zur naturkundlichen Forschung in Sudwestdeutschland, 26 (3), 137-169.
  • Wiltshire, E.P. (1980) The larger moths of Dhofar and their zoogeographic composition. Journal of Oman Studies, Special Report, 2, 187-216.
  • Wiltshire, E.P. (1990) An Illustrated. Annotated Catalogue of the Macro-Heterocera of Saudi Arabia. Fauna of Saudi Arabia, 11, 91-250.
  • Zullich, R. (1936) Beitrag zur Macrolepidopterenfauna des Rilogebirges in Bulgarien. Zeitschrift des Osterreichischen Entomologischen Vereines Wien, 21 (7/8), 49-55.